Gnathymenus penicillatus, Assing, 2015

Assing, Volker, 2015, New species and additional records of Dolicaonina from Ecuador (Coleoptera: Staphylinidae: Paederinae), Linzer biologische Beiträge 47 (2), pp. 1119-1132: 1120-1121

publication ID

persistent identifier

treatment provided by


scientific name

Gnathymenus penicillatus


Gnathymenus penicillatus   nov.sp. ( Figs 1-9 View Figs 1-9 )

T y p e m a t e r i a l: Holotype 3: " Ecuador Cotopaxi, Volcán Cotopaxi m 3925, 3.VIII.2006, P.M. Giachino / Holotypus 3 Gnathymenus penicillatus   sp. n. det. V. Assing 2014" ( SMNS)   . Paratypes: 733, 8♀♀: same data as holotype ( SMNS, cAss)   ; 2♀♀: " Ecuador Cotopaxi, Volcán Cotopaxi m 4000, 3.VIII.2006, Luca Picciau leg." ( SMNS)   .

E t y m o l o g y: The specific epithet is an adjective derived from the Latin noun penicillus (brush) and alludes to the pair of tufts of setae on the male sternite VII.

D e s c r i p t i o n: Body length 4.0- 4.6 mm; length of forebody 1.9-2.0 mm. Coloration: head reddish; pronotum reddish-brown to blackish-brown, at least slightly darker than head; elytra blackish; abdomen blackish with the apical segments (VIII-X) palereddish; legs reddish-brown to dark-brown with reddish tarsi; antennae reddish; maxillary palpi reddish with penultimate palpomere infuscate.

Head ( Fig. 1 View Figs 1-9 ) transverse, approximately 1.1 times as broad as long, lateral margins behind eyes subparallel in dorsal view; dorsal surface with sparse and moderately coarse punctation; median dorsal portion impunctate or with few punctures; interstices without microsculpture. Eyes small and weakly convex, approximately half as long as postocular region in dorsal view. Antenna approximately 1.0 mm long.

Pronotum ( Fig. 1 View Figs 1-9 ) approximately 1.15 times as long as broad and slightly broader than head, weakly tapering posteriad; midline broadly impunctate, laterally not delimited by a distinct dorsal series of punctures; punctation of lateral portions similar to that of head; interstices without microsculpture.

Elytra ( Fig. 1 View Figs 1-9 ) short, approximately 0.55 times as long as pronotum and with weakly marked humeral angles; punctation sparse, fine and shallow; interstices without microsculpture. Hind wings completely reduced. Metatarsomere I noticeably longer than II.

Abdomen broader than elytra; segments III-VI with distinct paratergites; punctation moderately sparse, finer on posterior than on anterior tergites; interstices with distinct transverse microsculpture; posterior margin of tergite VII without palisade fringe; posterior margin of tergite VIII obtusely angled in the middle ( Fig. 4 View Figs 1-9 ).

3: sternites III-V unmodified; sternite VI slightly flattened in the middle, otherwise unmodified; sternite VII ( Figs 2-3, 5-6 View Figs 1-9 ) strongly modified, with deep posterior excision, the surroundings of this excision deeply impressed, postero-laterally with two pronounced spine-like processes on either side, margin of posterior excision with a brush-like tuft of long setae on either side; sternite VIII ( Fig. 7 View Figs 1-9 ) oblong, 1.20-1.25 times as long as broad, with conspicuously deep median incision, this incision nearly reaching anterior margin of sternite; aedeagus ( Figs 8-9 View Figs 1-9 ) approximately 0.7 mm long and of distinctive shape; parameres completely reduced.

C o m p a r a t i v e n o t e s: This species is distinguished from all its congeners particularly by the distinctive morphology of the aedeagus and by the distinctive shape and chaetotaxy of the male sternite VII. Based on the similarly modified male sternite VII, G. penicillatus   is closely allied to G. praenuptus ROUGEMONT, 2014   from Ecuador (Imbabura), from which it is additionally separated by smaller body size ( G. praenuptus   : 6.2 mm). For illustrations of G. praenuptus   see ROUGEMONT (2014).

D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: The type locality is situated in the northeast of Cotopaxi province, Central Ecuador. The type specimens were sifted from litter in the so-called "Paramo formation" at altitudes of 3925 and 4000 m, near the summit of Cotopaxi Volcano (GIACHINO, pers. comm.).


Tavera, Department of Geology and Geophysics


Royal British Columbia Museum - Herbarium


Staatliches Museum fuer Naturkund Stuttgart