Anchista nubila Andrewes, 1931

Shi, Hongliang, Zhou, Hongzhang & Liang, Hongbin, 2013, Taxonomic synopsis of the subtribe Physoderina (Coleoptera, Carabidae, Lebiini), with species revisions of eight genera, ZooKeys 284, pp. 1-129 : 28-29

Anchista nubila Andrewes, 1931 Habitus: Figs 8 42 Habitus: male genitalia: Fig. 73

Anchista nubila Andrewes 1931: 526 (original: Anchista ; type locality: Dehra Dun; lectotype deposited in NHML); Csiki 1932: 1456 ( Anchista , catalogue).

Type examined.

Lectotype of Anchista nubila Andrewes, designated herein (NHML): female, body length = 8.5 mm, board mounted, "Dehra Dun / Rajpur, 3404 ft."; "Bought from / Staudinger & / Bang-Haas, 1930"; “Type” [red label]; " Anchista / nubila / Type Andr. / H. E. Andrewes det."; "H. E. Andrewes Coll. / B. M. 1945-97"; "LECTOTYPE ♀ / Anchista nubila / Andrewes, 1931 / des. SHI H. L. 2011" [Fig. 42]. Paralectotypes of Anchista nubila Andrewes: 1 male and 2 females (NHML), "Dehra Dun / Rajpur, 3404 ft."; "Bought from / Staudinger & / Bang-Haas, 1930"; “Cotype” [round label, with green circle]; "H. E. Andrewes Coll. / B. M.1945-97"; "PARALECTOTYPE / Anchista nubila / Andrewes, 1931 / det. SHI H. L. 2011".

Notes on types.

The original description mentioned five specimens from Dehra Dun in Andrewes’ collection, but did not designate a holotype. In the collection of NHML, we found only four specimens. Among them, a female specimen bears a determination label of Andrewes and a red label “Type”, and this is herein designated as lectotype ( Fig. 42 ) for taxonomic purpose of fixing the name to unique name-bearing type. The other three specimens have only a round label “co-type”, but no Andrewes’ determination label.

Non-type material examined

(Total 16 specimens from India). 9 specimens (MNHN), "Chota Nagpore, Nowatoli, R. P. Cardon, VIII-XI 1896"[Fig. 73]. 3 specimens (MNHN), "Chota Nagpore, Nowatoli, R. P. Cardon, V.VI.1896". 1 specimen (MNHN), "Inde Angl, Ghates, R. P. F. Tabourel, VII-IX 1898". 1 specimen (MNHN), "S. India, Nedungadu, P. S. Nathan, 1936"[Fig. 8]. 1 specimen (MNHN), "Chota Nagpore"; "Ex Musaeo, H. W. Bates, 1892". 2 specimens (NHML), "India mer. Tanjore Distr. Nedungadu".

Diagnosis.

Head and pronotum glabrous; pronotum widest near middle, lateral margins distinctly angulate at middle; elytra uniform brownish; 3rd interval of elytra with three or four pores, 5th interval with two or three pores near base.

The glabrous surface and the number of setigerous pores on the 3rd and 5th intervals readily distinguish this species from most allied species, except Anchista fenestrata fenestrata . Diagnosis between these two species is presented in the key. In general appearance, this species looks like Anchista brunnea , but can be distinguished from the latter by the wider and more angulate pronotum, and the different number of elytral setigerous pores.

Description.

Body length 7.5-8.5 mm; dorsal side uniform reddish brown, pronotum lateral explanate areas somewhat paler; elytra usually with margins darker, but not forming central patch; antennae reddish brown, 1st antennomere paler; femora yellowish, tibiae and tarsi darker; ventral side reddish brown. Head glabrous, without or with very fine isodiametric microsculpture. Pronotum glabrous,cordiform, widest slightly before middle; ratio PW/PL 1.30 to 1.41; pronotal base briefly but distinctly lobed; disc moderately convex, microsculpture indistinct, without punctures; front angles with few short setae (distinctly shorter than in Anchista fenestrata fenestrata ); lateral margins distinctly angulate in middle, strongly sinuate before hind angles; lateral explanate areas wide and flat, with few coarse punctures; hind angles sharp, rectangular or slightly acute, with few very short accessory setae; basal foveae moderately deep, with few punctures; median line distinct, strongly depressed and punctate anteriorly and subbasally. Elytra with striae shallow, distinct, finely punctate; intervals slightly convex, without accessory setae, finely and sparsely punctate; microsculpture distinct, isodiametric; 3rd interval usually with three setigerous pores, sometimes with four, all adjacent to 2nd interval, basal one usually placed at basal one-fourth approximately; 5th interval usually with two setigerous pores, sometimes with three, basal one near to interval base, all pores adjacent to 5th stria; basal margination nearly complete. Males with adhesive hairs rudimentary (two rows weakly present near apex) on 1st mesotarsomere, well developed (two whole rows) on 2nd and 3rd mesotarsomeres. Male genitalia with median lobe of aedeagus stout, bent to right side near apex; in lateral view, distinctly expanded in middle, gradually narrowed to apex; left lateral margin not curved in middle; apical orifice opened dorsally, basal margin of apical orifice with long setae; apical lamella placed on right side, flat, wide and rounded in dorsal view, not curved up dorsally; internal sac with a weakly sclerotized sclerite near middle, two weakly sclerotized elongate and sinuate sclerites near apical orifice, some parts of internal sac irregularly scaled ( Fig. 73 ). Female genitalia not studied.

Distribution

( Map 3 ). India.

Figures

Map 3. Known distribution of genus Anchista: ◆ Anchista brunnea (Wiedemann) ★ Anchista nubila Andrewes ● Anchista pilosa sp. n. △ Anchista fenestrata fenestrata (Schmidt-Goebel) ▼ Anchista fenestrata subpubescens Chaudoir.

Figure 73. Median lobe of male genitalia, right-lateral, ventral, left-lateral, dorsal views of Anchista nubila Andrewes, a specimen from Chota Nagpore.

Figures 7 - 12. Habitus, scale bars = 2.0 mm: 7 Anchista fenestrata subpubescens Chaudoir, a specimen from Pakistan 8 Anchista nubila Andrewes, a specimen from India 9 Metallanchista perlaeta (Kirschenhofer), a specimen from Sumatra 10 Metallanchista perlaeta (Kirschenhofer), a specimen from Java 11 Physodera dejeani Eschscholtz, a specimen from Yunnan 12 Physodera eschscholtzii Parry, a specimen from Yunnan.

Figures 37 - 42. Type materials, scale bars = 2.0 mm: 37 Lectotype of Lebia brunnea Wiedemann [= Anchista brunnea (Wiedemann)] 38 Lectotype of Anchista picea Chaudoir [= Anchista brunnea (Wiedemann)] 39 Holotype of Plochionus fenestratus Schmidt-Goebel [= Anchista fenestrata fenestrata (Schmidt-Goebel)] 40 Holotype of Anchista glabra Chaudoir [= Anchista fenestrata fenestrata (Schmidt-Goebel)] 41 Lectotype of Anchista subpubescens Chaudoir [= Anchista fenestrata subpubescens Chaudoir] 42 Lectotype of Anchista nubila Andrewes.

Figures 43 - 48. Type materials, scale bars = 2.0 mm: 43 Holotype of Anchista pilosa sp. n. 44 Holotype of Metallanchista laticollis sp. n. 45 Holotype of Diamella kaszabi Jedlicka [= Diamella kaszabi (Jedlicka)] 46 Lectotype of Calleida cupreomicans Oberthuer [= Diamella cupreomicans (Oberthuer)] 47 Lectotype of Allocota aerata Bates [= Diamella cupreomicans (Oberthuer)] 48 Holotype of Lachnoderma arrowi Jedlicka [= Diamella arrowi (Jedlicka)].