Cunninghamites squamosus Heer, 1871

Halamski, Adam T., 2013, Latest Cretaceous leaf floras from southern Poland and western Ukraine, Acta Palaeontologica Polonica 58 (2), pp. 407-443 : 412-414

Cunninghamites squamosus Heer, 1871

Figs. 4 , 5A–C.

1871 Cunninghamites squamosus m.; Heer 1871: 9–10, pl. 1: 5–7.

1907 Cunninghamia elegans (Corda) Endl. ; Nowak 1907a: 47, pl. 1: 11.

1962 Cunninghamia elegans (Corda) Endl. ; Ciesliński and Milaković 1962: 253, pl. 1: 3.

1967 Cunninghamia elegans Corda ; Malicki et al. 1967: 226–227, fig. 17.

1967 Cunninghamia stenophylla Velen. ; Malicki et al. 1967: 227, figs. 18–19.

1975 Cunninghamia stenophylla Velen. ; Kohlman−Adamska 1975: 163, 164, pl. 1: 6.

2009 Cunninghamites squamosus Heer 1871 ; Bosma et al. 2009: 486– 487, figs. 3A, 4C–F, 5.

2012 Cunninghamites squamosus Heer ; Bosma et al. 2012: 24–25, pls. 1: 5, 6; 2: 4, 7 [ubi syn.].

Type material: lectotype, fragmentary twig BSPG coll. Univ. Würzburg no. 6166 (ex coll. Schenk), designated by Bosma et al. (2012: 24); see Bosma et al. (2012: 24) for the list of epitypes.

Type locality: “Altenburg” near Quedlinburg , Saxony−Anhalt, Germany .

Type horizon: Heidelberg Formation , upper Santonian .

Material.—Fragmentary leafy twigs: two from Chełm, MZ VII/32/13 (investigated under the SEM) and MZ VII/33/1; single specimen from Jasionna, PIG 60 III 3 and from Potelych, L PB−K.16 (formerly 7678); two from Krasnobród, MZ VII/32/11 and JS 6.

Description.—The preserved part of shoots up to 10 cm long; leaf scars rhomboid, ca. 3 mm wide and 3.5–5 mm long. Leaves spirally arranged, either straight or slightly adaxially incurved, linear (Chełm) or lanceolate (other localhttp://dx.doi.org/10.4202/app.2011.0024

Type locality: Kunrade , Limburg, the Netherlands .

Type horizon: Maastricht Formation , upper Maastrichtian .

Material.—Fragmentary leafy twigs MZ VII/33/36 from Krasnobród and ZPAL Pl 4/9/1 from Wola Pawłowska.

Description.—Axis fragment unbranched, ca. 10 cm long. Leaf base cushions poorly preserved. Leaves decurrent, their free portions spreading in all directions at an angle not exceeding 30 °, dorsiventrally flattened, linear, up to at least 9 cm long (apices not observed) and 2–4 mm wide. Margin entire. Blade with a distinct median keel or vein, sometimes also with finer longitudinal striations.

ities), up to at least 40 mm long and 2 mm wide; the greatest width is in the basal part.

Leaves of the Chełm specimens ca. 1.2 mm wide ( Fig. 4 ). Two parallel stomatal bands ca. 350 µm wide separated by an intercostal zone ca. 300 µm wide; intercostal cells elongate, ca. 20 µm wide.

Remarks.—The cuticular features of the specimen MZ VII/ 32/13 from Chełm, although poorly preserved, allow to refer the described material to Cunninghamites squamosus Heer, 1871 , intitally described from the Santonian of Quedlinburg, characterised by absence of microscopic marginal serration ( Bosma et al. 2009). C. lignitum from the Cenomanian of Bohemia has serrate leaves ( Kvaček 1999).

The material from Chełm is rather different in overall aspect when compared to specimens from other localities, the leaves being narrower and linear. This feature is tentatively interpreted as reflecting intraspecific variability and preservation artifacts (chalk versus “opoka”) rather than taxonomically important characters. Stratigraphic and geographic range.—North America and

Europe, Santonian to Maastrichtian ( Bosma et al. 2012).

Cunninghamites ubaghsii Debey ex Ubaghs, 1885 emend. van der Ham , van Konijnenburg−van Cittert, and Nieuwenhuis, 2004

Figs. 5J, 6B.

1885 Cunninghamites Ubaghsi ; Ubaghs 1885: 28.

2004 Cunninghamites ubaghsii Debey ex Ubaghs, 1885 ; van der Ham et al. 2004: 89–94, figs. 3–6 [ubi syn.].

2012 Cunninghamites ubaghsii Debey ex Ubaghs ; Bosma et al. 2012: 25; pl. 2: 1.

Type material: holotype, subcomplete twig IRSNB−Paleobot. b4318

(van der Ham et al. 2004: fig. 3).

Remarks.—Two relatively large bark impressions devoid of leaves (therefore specifically unidentifiable) from Krasnobród (KrM Pb 2/1, 2) are referred here as Cunninghamites sp. ( Fig. 5I). Leaf cushions are obovoid−rhomboid, spirally arranged, 8–10 mm long and 5–7 mm wide.

Stratigraphic and geographic range.— Limburg (Kunrade and vicinity of Maastricht), upper Maastrichtian (van der Ham et al. 2004); Roztocze (Krasnobród and Wola Pawłowska), uppermost Campanian.

Figures

Fig. 4. Conifer Cunninghamites squamosus Heer, 1873. Single leaf from the specimen MZ VII/32/13, Chełm, Maastrichtian, SEM view.