Cicindela (s. str.) herbacea herbacea Klug, 1832

Matalin, Andrey V. & Chikatunov, Vladimir I., 2016, The tiger beetles (Coleoptera, Carabidae, Cicindelinae) of Israel and adjacent lands, ZooKeys 578, pp. 115-160 : 121

(○) Cicindela (s. str.) herbacea herbacea Klug, 1832

General distribution.

Asia Lebanon.

References.

Lebanon - Wiesner 1992: 127; Puchkov and Matalin 2003: 105; Franzen 2007: 13; Deuve 2011: 129.

Distribution.

Lebanon, Liban-Nord: Bcharré, Les Cèdres, VI. 1997 1♂1♀ (cIOv); Bcharré 1♀ (after Franzen 2007); Tannourine 1♀ (after Deuve 2011).

Comments.

Until recently both these species were recorded from Syria, Lebanon and Israel by several authors as Cicindela herbacea Klug ( Valdenberg 1983; Nussbaum 1987; Wiesner 1992; Puchkov and Matalin 2003; Chikatunov et al. 2006; Franzen 2007; Ptashkovsky 2009). However, according to recent data Cicindela herbacea does not occur in Israel ( Deuve 2011, 2012). The nominative subspecies occurs in Lebanon and Syria; Cicindela herbacea aleppensis Deuve, 2012 is recorded from north-western Syria, while Cicindela herbacea perreaui Deuve, 1987 and Cicindela herbacea colasi Deuve, 2011 are found in Turkey Tunceli and Adana Provinces, respectively. On the basis of the shape of pronotum (Figs 38 vs 39), white elytral pattern (Figs 54 vs 55), size of aedeagus and shape of it apex (Figs 93 vs 97), as well as shape of internal sack (Figs 94-96 vs 98-100) we consider all studied specimens from Israel to be Cicindela javeti azari Deuve, 2011 (type locality - Lebanon, Jezzine). It should be noted that the taxonomy of intraspecific forms within the ' campestris '-group is complex, and additional studies are necessary.

Figures

Figures 30 - 45. Pronotum of males, dorsal view: 30 Calomera aulica aulica 31 Calomera littoralis aulicoides 32 Calomera littoralis winkleri 33 Calomera littoralis nemoralis 34 Cephalota tibialis tibialis 35 Cephalota littorea littorea 36 Cephalota zarudniana vartianorum 37 Cephalota deserticola deserticola 38 Cicindela herbacea herbacea 39 Cicindela javeti azari 40 Habrodera nilotica nilotica 41 Lophyra flexuosa flexuosa 42 Hypaetha singularis 43 Cylindera contorta valdenbergi 44 Myriochila melancholica melancholica 45 Grammognatha euphratica euphratica; pr st pronotopleural suture. Scale bars: 1 mm (30 - 33: A; 34 - 37: B; 38 - 41: C; 42 - 43: D; 44: E; 45: F).

Figures 46 - 61. Left elytron of males, dorsal view: 46 Calomera aulica aulica 47 Calomera littoralis aulicoides 48 Calomera littoralis winkleri; 49 Calomera littoralis nemoralis 50 Cephalota tibialis tibialis 51 Cephalota littorea littorea 52 Cephalota zarudniana vartianorum 53 Cephalota deserticola deserticola 54 Cicindela herbacea herbacea 55 Cicindela javeti azari 56 Habrodera nilotica nilotica 57 Lophyra flexuosa flexuosa 58 Hypaetha singularis 59 Cylindera contorta valdenbergi 60 Myriochila melancholica melancholica 61 Grammognatha euphratica euphratica. Scale bars: 1 mm.

Figures 93 - 100. Aedeagus and internal sack of Cicindela ssp.: 93 - 96 Cicindela herbacea herbacea 97 - 100 Cicindela javeti azari 93, 97 aedeagus 94 - 96, 98 - 100 internal sac 93, 96 - 97, 100 left lateral view 95, 99 dorsal view 94, 98 right lateral view 98 - 100 partly inflanted); B basal bladder; VLR - ventro-lateral right bladder. Scale bars: 1 mm (93, 97: A; 94 - 96: B; 98 - 100: C).