Himalayana martensi Grismado, 2014

Grismado, Cristian J., Deeleman, Christa, Piacentini, Luis N., Izquierdo, Matías A. & Ramírez, Martín J., 2014, Taxonomic Review Of The Goblin Spiders Of The Genus Dysderoides Fage And Their Himalayan Relatives Of The Genera Trilacuna Tong And Li And Himalayana, New Genus (Araneae: Oonopidae), Bulletin of the American Museum of Natural History 2014 (387), pp. 1-108 : 77-82

Himalayana martensi Grismado , new species

Figures 66–68 , 72C–D

TYPES: Male holotype , one male and one female paratypes from Nepal: Gandaki: Manang Dist., Marsyandi, 2200 m, Apr. 12–13, 1980, Martens and Ausobsky ( SGN PBI_OON 14923 ).

ETYMOLOGY: The specific name is a noun in apposition in honor to Jochen Martens, one of the collectors of this and other oonopid species from the Himalayas.

DIAGNOSIS: Males of this species are recognizable by the following combination of characters: copulatory bulb with globose basal and median part, slightly tickened palpal femur (figs. 68C–E, 72D) and stout, forwardly directed enlarged setae on the posterior part of the epigastric scutum (fig. 68B); in the females, the internal genitalia resemble those of H. kathmandu , but the lateral apodemes are shorter (fig. 72C). In addition, both sexes are smaller than H. kathmandu (total length <1.8 mm) and only slightly sclerotized.

DESCRIPTION: Male (holotype, PBI_OON 14923). Total length 1.64. Cephalothorax: Carapace orange, pars cephalica slightly elevated in lateral view, anteriorly narrowed to 0.49 times its maximum width or less, sides granulate; lateral margin rebordered; nonmarginal pars cephalica setae needlelike, in U-shaped row; nonmarginal pars thoracica setae needlelike; marginal setae needlelike. Eyes ALE largest, PME circular, PLE circular; posterior eye row recurved from above, straight from front; ALE separated by their radius to diameter, ALE–PLE separated by less than ALE radius, PME separated by less than their radius. Sternum longer than wide, lateral margins unmodified, orange; setae evenly scattered. Mouthparts: chelicerae, labium, and endites orange; fangs directed medially. Abdomen: Dorsum soft portions not visible from above. Book lung covers elliptical. Pedicel tube short. Dorsal scutum orange. Epigastric scutum not protruding, orange. Postepigastric scutum orange. Spinneret scutum with fringe of needlelike setae. Colulus present. Legs: Pale orange. Leg spination (all spines longer than segment width): leg I: tibiae v2-2-2-0-0, metatarsi v2- 2-0; leg II: tibiae v2-2-2-0-0, metatarsi v2-2-0. Tarsi I–IV superior claw tooth not examined in detail. Trichobothria not examined. Genitalia: Epigastric region with sperm pore small, oval, unmodified; stout, forwardly directed enlarged setae on the posterior part of the epigastric scutum (fig. 68B). Palpal embolus with one dorsal and one prolateral accompanying laminae with filiform projections (more numerous in the latter, fig. 72D); femur slightly thickened; patella nearly as long as tibia; cymbium narrow in dorsal view; bulb pale orange, distal part with a dorso-prolateral pointed, short, acute projection (fig. 72D).

Female (paratype, PBI_OON 14923). Total length 1.74. As in male except as noted. Cephalothorax: Eyes PLE–PME touching. Palp without spines. Abdomen: Dorsal scutum covering more than 3/4 of abdomen, orange. Epigastric scutum not strongly sclerotized, orange. Postepigastric scutum not strongly sclerotized, not fused to epigastric scutum, orange. Legs: Patella plus tibia I shorter than carapace. Leg spination (all spines longer than segment width): leg I: tibiae: v2-2-2-2-0, metatarsi v2-2-0; leg II: tibiae v2-2-2-0-0, metatarsi: v2-2-0. Genitalia: Copulatory opening rounded, large, visible with stereomicroscope (fig. 66H). Lateral apodemes short (fig. 72C).

OTHER MATERIAL EXAMINED: None.

DISTRIBUTION: Known only from the type locality.

Figures

Fig. 66. Himalayana martensi, new species, female (PBI_OON 14923). A. Habitus, dorsal view. B. Same, ventral view. C. Same, lateral view. D. Carapace, dorsal view. E. Same, ventral view. F. Same, lateral view. G. Same, anterior view. H. Epigastric area, ventral view.

Fig. 67. Himalayana martensi, new species, male (PBI_OON 14923). A. Habitus, dorsal view. B. Same, ventral view. C. Same, lateral view. D. Carapace, dorsal view. E. Same, ventral view. F. Same, lateral view.

Fig. 68. Himalayana martensi, new species, male (PBI_OON 14923). A. Carapace, anterior view. B. Abdomen, ventral view. C. Left palp, retrolateral view. D. Same, dorsal view. E. Same, prolateral view.

Fig. 72. Himalayana spp. genitalia. A, B. Himalayana kathmandu, new species. A. Female (PBI_OON 15258), internal genitalia, dorsal view. AP = apodeme; AS = anterior sclerite; PR = posterior receptacle; TP = transverse plates; arrow = glandular field. B. Male (PBI_OON 15268), left palp, retrolateral view. Arrow = acute dorsal projection; GT = glandular tube). C, D. Himalayana martensi, new species. C. Female (PBI_OON 14923), internal genitalia, dorsal view. D. Male (PBI_OON 14923), left palp, prolateral view. E, F. Himalayana castanopsis, new species. E. female (PBI_OON 15747), internal genitalia, dorsal view. F. Male (PBI_OON 15978), left palp, prolateral view.