Camptoscaphiella glenniei ( Fage, 1946 ) Grismado & Deeleman & Piacentini & Izquierdo & Ramírez, 2014

Grismado, Cristian J., Deeleman, Christa, Piacentini, Luis N., Izquierdo, Matías A. & Ramírez, Martín J., 2014, Taxonomic Review Of The Goblin Spiders Of The Genus Dysderoides Fage And Their Himalayan Relatives Of The Genera Trilacuna Tong And Li And Himalayana, New Genus (Araneae: Oonopidae), Bulletin of the American Museum of Natural History 2014 (387), pp. 1-108 : 104-106

Camptoscaphiella glenniei ( Fage, 1946) , new combination

Figure 83

Triaeris glenniei Fage 1946: 384

(female holotype from India: Uttarakhand: Dehra Dun: Chakrata Taksil: Moila cave , 2691 m, N 30.77694 °, E 77.78722 °, 12–19.I.1946, Brig. E.A. Glennie, deposited in BMNH, PBI_OON 43075 ; examined).

NOTES: The heart-shaped sternum, the general morphology of the carapace, the size of the abdominal scuta, the leg spine pattern, and, especially, the genitalic conformation show no significant differences from all the other known species of Camptoscaphiella (see Baehr and Ubick, 2010). The slightly longer legs and the reduced eyes of this species are here considered as adaptations to the cavernicolous habitat rather than as an indication for a different generic placement.

Given that the holotype is the only known specimen, we decided not to dissect the genitalia for fine study of the internal details (as, for example, the disposition of apodemes). We opted only to clarify the entire abdomen to assign its generic placement temporarily and to obtain a gross differentiation from the other known species (fig. 83H–J).

DIAGNOSIS: Camptoscaphiella glenniei differs from all congeners by its reduced eyes (fig. 83D, G). Camptoscaphiella sinensis Deeleman-Reinhold, 1995 , is a completely eyeless species, while all the other Camptoscaphiella have normally developed eyes ( Baehr and Ubick, 2010).

REDESCRIPTION: Female (holotype): Total length 2.65. Cephalothorax: Carapace pale orange, without any pattern, broadly oval in dorsal view, pars cephalica strongly elevated in lateral view, anteriorly narrowed to 0.49 times its maximum width or less, with rounded posterolateral corners, posterolateral edge without pits, posterior margin not bulging below posterior rim, posterolateral surface without spikes, surface of elevated portion of pars cephalica smooth, sides finely reticulate, thorax without depressions, fovea absent, without radiating rows of pits; lateral margin undulate, rebordered, without denticles; plumose setae near posterior margin of pars thoracica absent; nonmarginal pars cephalica setae light, needlelike, scattered; nonmarginal pars thoracica setae absent; marginal setae absent. Clypeus margin unmodified, straight in front view (fig. 83G), sloping forward in lateral view (fig. 83F), high, ALE separated from edge of carapace by their radius or more, median projection absent; setae light, needlelike. Chilum absent. Eyes six, reduced (but apparently still functional, fig. 83D), ALE largest, ALE oval, PME circular, PLE circular; posterior eye row straight from above, procurved from front; ALE separated by their radius to diameter, ALE–PLE separated by less than ALE radius, PME touching for less than half their length, PLE–PME separated by PME radius to PME diameter. Sternum as long as wide, pale orange, uniform, not fused to carapace, median concavity absent, without radial furrows between coxae I–II, II–III, III–IV, surface smooth, without pits, microsculpture absent, sickle-shaped structures absent, anterior margin unmodified, posterior margin not extending posteriorly of coxae IV, lateral margin without infracoxal grooves, distance between coxae approximately equal, extensions of precoxal triangles present, lateral margins unmodified, without posterior hump; setae sparse, light, needlelike, evenly scattered, originating from surface. Mouthparts: chelicerae, endites, and labium pale orange. Chelicerae straight, anterior face unmodified; directed medially, shape normal, without prominent basal process, fang tip unmodified; setae light, needlelike, evenly scattered; paturon distal region unmodified, posterior surface unmodified, promargin unmodified, inner margin unmodified. Labium triangular, not fused to sternum, anterior margin indented at middle. Endites distally not excavated, serrula present in single row, anteromedian tip unmodified, posteromedian part unmodified. Palp spines and claws absent; tarsus unmodified. Abdomen: Ovoid, without long posterior extension, rounded posteriorly, interscutal membrane rows of small sclerotized platelets absent; dorsum soft portions pale white, without color pattern. Book lung covers large, ovoid, without setae, anterolateral edge unmodified. Posterior spiracles connected by groove. Pedicel tube short, ribbed, scutopedicel region unmodified, scutum not extending far dorsal of pedicel, plumose hairs absent, matted setae on anterior ventral abdomen in pedicel area absent, cuticular outgrowths near pedicel absent. Dorsal scutum weakly sclerotized, pale orange, without color pattern, covering less than 1/2 of abdomen, between 1/4 and 1/2 abdomen width, not fused to epigastric scutum (fig. 83G), middle surface and sides smooth, anterior half without projecting denticles. Epigastric scutum weakly sclerotized, surrounding pedicel, not protruding, small lateral sclerites absent. Postepigastric scutum weakly sclerotized, pale orange, widely hexagonal, only around epigastric furrow, not fused to epigastric scutum, anterior margin with triangular lateral joints fitting under epigastic scutum. Spinneret scutum absent. Supraanal scutum absent. Dorsum setae light, needlelike. Epigastric area setae uniform, light, needlelike. Postepigastric area setae, light, needlelike. Dense patch of setae anterior to spinnerets absent. Colulus represented only by setae. Legs: Pale orange, without color pattern; femur IV not thickened, same size as femora I–III, patella plus tibia I longer than carapace, tibia I unmodified. Leg spination (all spines longer than segment width): leg I: femur: pv0-0-1-1-0-0; tibia: v2-2-2-2-0; metatarsus: v2-2-0, leg II: tibia: v2-2-2-2-0; metatarsus: v2-2-0, leg III: tibia: v1ap, leg IV: tibia: v1ap. Tarsi I–IV superior claws tooth not examined in detail. Tarsi I to IV without inferior claw. Trichobothria not examined. Genitalia: External copulatory opening slitlike (fig. 83H–J). Copulatory duct narrow, long, with its proximal half directed posteriorly, and distal half sinuous, turned to dorsal. The area where the duct is bent looks (in ventral view) as a darkened thickening (fig. 83I).

Male: Unknown.

OTHER MATERIAL EXAMINED: None.

DISTRIBUTION: Known only from the type locality.

Figures

Fig. 83. Camptoscaphiella glenniei (Fage), new combination, female holotype (PBI_OON 43075). A. Habitus, dorsal view. B. Same, ventral view. C. Same, lateral view. D. Carapace, dorsal view. E. Same, ventral view. F. Same, lateral view. G. Habitus, anterior view. H. Epigastric area, ventral view. I. Same, cleared, ventral view. J. Same, lateral view.