Glenognatha patriceae, Cabra-García & Brescovit, 2016

Jimmy Cabra-García & Antonio D. Brescovit, 2016, Revision and phylogenetic analysis of the orb-weaving spider genus Glenognatha Simon, 1887 (Araneae, Tetragnathidae), Zootaxa 4069 (1), pp. 1-183 : 72-78

publication ID

https://doi.org/10.11646/zootaxa.4069.1.1

publication LSID

lsid:zoobank.org:pub:52FC658C-78C7-49FC-9961-8AC43CA03101

DOI

https://doi.org/10.5281/zenodo.5666946

persistent identifier

https://treatment.plazi.org/id/8E0E8F67-7D55-FFC5-ACBA-0E5DFCD1D272

treatment provided by

Plazi

scientific name

Glenognatha patriceae
status

new species

Glenognatha patriceae new species

( Figs. 56 View FIGURE 56 –59, 132)

Types. Male holotype from Hacienda San Pedro, Alto Queremal , Dagua , Valle del Cauca, Colombia, 3º29´N, 76º42´W, elev. 1800m, 21.ix.2009, J. Cabra leg., deposited in ICN 5987 View Materials GoogleMaps ; Paratypes: same data as the holotype, 4♂ 19♀ deposited in ICN 5986 View Materials GoogleMaps ; 1♂ 1♀ deposited in IBSP 163858.

Etymology. The species epithet is in memory of the first author’s mother, Patricia García Pinilla. The name is a noun in genitive.

Diagnosis. Males of G. patriceae resemble those of G. gaujoni and G. globosa by the presence of thin pleural bars between coxae II–III ( Fig. 56C View FIGURE 56 ) and by the slender palpal tibia (longer than 1.5 times the maximum tibia width) ( Figs. 58 A, E View FIGURE 58 ). It can be distinguished from the former by the absence of scale-like projections in the embolus ( Figs. 58D View FIGURE 58 , 59B–C) and from the latter by the thin distal projection of the conductor lamina ( Figs. 58B–C View FIGURE 58 , 59B–C). Females can be easily distinguished from all other Glenognatha species (except G. gaujoni ) by the presence of cheliceral ectal apophysis ( Figs. 57D–F View FIGURE 57 ); distinguished from G. gaujoni by the absence of anterior tooth ( Fig. 57D View FIGURE 57 ).

Male (holotype). Habitus as in Figures 56 A –D View FIGURE 56 . Carapace pale brown, darker along furrows. Sternum pale brown, darker on sides. Abdomen oval, dorsally with irregular pattern of two longitudinal dusky bands on sides. Silver guanine spots scattered in two median longitudinal bands and on lateral surfaces. Venter with a pale black median band and two lateral lighter bands with scattered silver guanine spots. Legs yellow-brownish. Tracheal spiracle well-sclerotized. Pleural bars between coxae I–II, II–III and III–IV. Total length 3.6. Cephalothorax 1.75 long, 1.30 wide, 0.90 high. Abdomen 2.15 long, 1.55 wide, 1.45 high. Sternum 0.90 long, 0.90 wide. Eye diameters and interdistances: AME 0.12. ALE 0.10. PME 0.15. PLE 0.10. AME-AME 0.10, AME-ALE 0.15, PME-PME 0.21, PME-PLE 0.12, ALE-PLE 0.01. Clypeus height 1.2. Chelicerae slightly lighter than the cephalothorax and anteriorly projected ( Fig. 56B View FIGURE 56 ). Six promarginal and eight retromarginal teeth. Ret1 on the posterior surface ( Figs. View FIGURE 57

57B–C). Leg measurements: femur 3.20, patella 0.75, tibia 3.25, metatarsus 3.35, tarsus 1.65, total 12.20; II: 3.00, 0.65, 3.00, 2.90, 1.40, 10.95; III: 2.00, 0.40, 1.30, 1.40, 0.55, 5.65; IV: 2.55, 0.50, 2.05, 2.10, 0.87, 8.07. DTS 0.50. DTE 0.65. Palp as in Figures 58E–H View FIGURE 58 . Palp tibia longer than wide ( Fig. 58 A, E View FIGURE 58 ). Conductor lamina distal end thin ( Figs. 58B–C View FIGURE 58 , 59B–C). Embolus tip nearly 180º curved ( Figs. 58D View FIGURE 58 , 59 A –C).

F IGURE 59. Glenognatha patriceae new species. A –C, male left palp. A, ventral. B, embolus and conductor, ventral. C, embolus and conductor, dorsal. D–G, female genitalia. D, dorsal. E, ventral. F, lateral. G, uterus externus distal portion. H–K, female tracheal system. H, dorsal, whole montage. I, dorsal. J, median and lateral trunks. K, median trunk transversal cut. Scale bars, 100 µm (A –F, H–J), 20 µm (G, K). C: conductor. CRa: conductor retrolateral apophysis. E: embolus. EMg: embolic medial groove. LT: lateral trunk. MC: membranous chamber. MT: median trunk. P: paracymbium. T: tegulum. UE: uterus externus.

Female (paratype ICN 5986). Coloration and general habitus as in male ( Figs. 56E–H View FIGURE 56 ). Pleural bars between coxae I–II, II–III and III–IV. Total length 4.8. Cephalothorax 2.05 long, 1.50 wide, 1.05 high. Abdomen 3.25 long, 2.55 wide, 2.45 high. Sternum 0.95 long, 0.90 wide. Eye diameters and interdistances: AME 0.10. ALE 0.10. PME 0.17. PLE 0.12. AME-AME 0.10, AME-ALE 0.12, PME-PME, 0.17 PME-PLE 0.15, ALE-PLE 0.02. Clypeus height 1.30. Chelicerae slightly lighter than the cephalothorax. Five promarginal and eight retromarginal teeth. Ectal apophysis well developed ( Figs. 57D–F View FIGURE 57 ). Leg measurements: I: femur 3.40, patella 0.70, tibia 3.45, metatarsus 3.35, tarsus 1.70, total 12.60; II: 3.25, 0.70, 3.05, 3.00, 1.40, 11.40; III: 1.85, 0.50, 1.55, 1.35, 0.55, 5.80; IV: 2.50, 0.50, 2.00, 2.10, 0.90, 8.00. DTS 0.70. DTE 0.55. Tracheal system as in Figures 59H–K. Genitalia without spermathecae (Figs. 59D–G). UE distal portion tubiform (Figs. 59D–G).

Variation. Male total length 3.9–4.3 (N =10; average 4.1), cephalothorax length 1.5–2.0 (N = 10; average 1.8), femur I length 3.2–3.7 (N = 10; average 3.4). Female total length 3.5–5.0 (N = 10; average 4.4), cephalothorax length 1.5–2.2 (N = 10; average 1.8), femur I length 3.0–4.0 (N = 10; average 3.4).

Distribution. Known from Valle del Cauca, Risaralda and Nariño states in Colombia ( Fig. 132 View FIGURE 132 ).

Additional material examined (N = 85). COLOMBIA: Quindio: Filandia , [4º40´N, 75º40´W], elev. [1758m], 4.viii.2012, N. Hazzi leg., 5♀ 1♂ 4 immature ( ICN 5998 View Materials ) GoogleMaps ; 2♀ 2♂ (IBSP 163859). Nariño: Barbacoas, Altaquer, Reserva Río Ñambi , [1º18´N, 78º5´W], elev. 1440m, 12.iv.2010, E. Flórez y Est. Tax. Animal UN leg., 2♀ 1♂ 1 immature ( ICN 4505 View Materials ) GoogleMaps ; 10.x.2009, E. Flórez y Est. Tax. Animal UN leg., 1♀ 2 immature (ICN 4408); 20.x.2012, N. Hazzi leg., 5♀ 1♂ 2 immature ( ICN 5999 View Materials ). Risaralda: Mistrato, San Antonio del Chambi , La Estrella Arcacay, [5º17´N, 75º53´W], elev. 900m GoogleMaps – 1100m, 31.iii–1.iv.1992?, Estud. Biol. UNAL leg., 1♀ (ICN 1078b). Valle del Cauca: Cali, El Palomar , [3º32´N, 76º36´W], elev. [1700m], 17.i.2013, J. Cabra & D. Rodríguez leg., 1♂ ( ICN 5994 View Materials ) GoogleMaps ; Cali, PNN Los Farallones , [3º15´N, 76º39´W], elev. 2050m, 2.iv.2000, M. M. Medina leg., 2♂ ( ICN 4317 View Materials ) GoogleMaps ; Cali , El Topacio, [3º19´N, 76º38´W], elev. [1624m], 6.i.2011, N. Hazzi leg., 1♀ 3♂ ( ICN 5996 View Materials ) GoogleMaps ; Dagua, same data as holotype, 37 immature ( ICN 5988 View Materials ); La Cumbre, Vereda Chicoral , 3º34´N, 76º35.6´W, elev. 1800m GoogleMaps – 2100m, ix.2009, J. Mendez leg., 1 immature (ICN 5989); Pance, ca. Topacio , [3º20´10.73´´N, 76º36´5.04´´W], elev. [1430m], viii.2011, N. Hazzi leg., 1♀ ( ICN 5990 View Materials ) GoogleMaps . Pance , 3º19´41´´N, 76º38´12´´W, elev. 1624m, 2.i.2012, N. Hazzi leg., 2♀ 3♂ 4 immatures ( ICN 5997 View Materials ) GoogleMaps .

DTE

Centro de Investigaciones Cient�ficas y Transferencia de Tecnolog�a a la Producci�n (CICyTTP-CONICET)

ICN

Instituto de Ciencias Naturales, Museo de Historia Natural

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Tetragnathidae

Genus

Glenognatha