Zeanillus punctigerus (Broun),

Sokolov, Igor M., 2016, A taxonomic review of the anilline genus Zeanillus Jeannel (Coleoptera: Carabidae: Bembidiini) of New Zealand, with descriptions of seven new species, re-classification of the species, and notes on their biogeography and evolution, Zootaxa 4196 (1), pp. 1-37: 11-16

publication ID


publication LSID


persistent identifier


treatment provided by


scientific name

Zeanillus punctigerus (Broun)


Zeanillus punctigerus (Broun) 

Figs. 1View FIGURE 1 B, 4B, 5B, 6B, 7B, 8A, 9B, 10BGM, 11B, 12A –D, 14B, 15B, 16B, 18A

Anillus punctigerus Broun, 1914: 153  . Type locality: « Mount Hutt , near Methven » (original citation). 

Examined material. Thirteen specimens (5 dissected) from the following localities: Canterbury: Staveley, Sharplin Falls ; Geraldine, Orari Gorge. 

Recognition. Adults of this species ( Fig. 11View FIGURE 11 B) are practically indistinguishable externally from the adults of Z. phyllobius  and Z. nunni  , and are distinguished from the males of those by the structure of male genitalia. From other congeners, adults of Z. punctigerus  can be distinguished by the characters listed in the subgeneric diagnosis.

Description. Size. Medium to large for genus (SBL range 1.64–1.94 mm, mean 1.80± 0.107 mm, n=5).

Habitus. Body form slightly convex, ovoid, moderately elongate (WE/SBL 0.35±0.010), head of moderate width for genus compared to pronotum (WH/WPm 0.76±0.016), pronotum of moderate width in comparison to elytra (WPm/WE 0.76±0.012).

Color. Body color rufotestaceous, appendages testaceous.

Prothorax. Pronotum moderately long (LP/LE 0.44±0.029) and moderately elongate (WPm/LP 1.29±0.032), with lateral margins rectilinear constricted posteriorly (WPm/WPp 1.70±0.118). Width between anterior angles much greater than between posterior angles (WPa/WPp 1.33±0.090).

Elytra. Slightly depressed along suture, comparatively short (LE/SBL 0.47±0.010) and moderately wide (WE/ LE 0.74±0.028). Lateral margins slightly divergent at basal third, subparallel at middle and evenly rounded to apex in apical third.

Male genitalia. Median lobe ( Fig. 12View FIGURE 12 A) with straight ventral margin and semicircular short apex. Apical orifice area with a pair of long ostial fields. Sclerites of internal sac united basally, long, slightly shorter than length between apical and basal orifices. Left paramere with apical constriction ( Fig. 12View FIGURE 12 C). Ring sclerite with long handle of approximately equal length and width ( Fig. 14View FIGURE 14 B).

Female internal genitalia. Goncoxite 2 with short ensiform setae ( Fig. 15View FIGURE 15 B). Spermatheca sclerotized, elongate, with three compartments ( Fig. 16View FIGURE 16 B). Spermathecal duct short with few coils. Attachments of spermathecal duct and gland to spermatheca close together.

Geographical distribution. This species is known from the eastern ranges of Canterbury lying in basins of the Rakaia, Ashburton and Rangitātā rivers ( Fig. 18View FIGURE 18 A, green circles). 

Way of life. Specimens were collected in the broadleaf forest.

Relationships. The structure of male genitalia and spermatheca suggests that Z. punctigerus  and the allopatric Z. phyllobius  from the coastal hills of the Banks Peninsula are the sister taxa.














Zeanillus punctigerus (Broun)

Sokolov, Igor M. 2016

Anillus punctigerus

Broun 1914: 153