Zeanillus carltoni, Sokolov, Igor M., 2016

Sokolov, Igor M., 2016, A taxonomic review of the anilline genus Zeanillus Jeannel (Coleoptera: Carabidae: Bembidiini) of New Zealand, with descriptions of seven new species, re-classification of the species, and notes on their biogeography and evolution, Zootaxa 4196 (1), pp. 1-37: 27

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Zeanillus carltoni


Zeanillus carltoni  , sp.n.

Figs. 11View FIGURE 11 G, 13A –D, 14J, 15G, 17A, 18B

Type material. HOLOTYPE, male, in NZAC, board-mounted, dissected, labeled: / New Zealand DN Mt Watkin Road at Mt Watkin 11 Aug 0 7 / Washed soil sample 420m tussockland /. 

PARATYPES (3 specimens, 2 dissected), 1 male and 2 females labeled same as holotype.

Specific epithet. The specific epithet honors the surname of Christopher E. Carlton, the Director of Louisiana State Arthropod Museum, well-known specialist in Pselaphinae  , for his magnanimous efforts of making the material on New Zealand Anillina available for study 

Type locality. New Zealand, South Island, Otago, Mount Watkin   .

Recognition. Adults of this species ( Fig. 11View FIGURE 11 G) are distinguished from other congeners by the characters listed in the species key.

Description. Size. Large for genus (SBL range 1.89–2.01 mm, mean 1.96± 0.050 mm, n=4).

Habitus. Body form slightly convex, slightly ovoid, moderately elongate (WE/SBL 0.35±0.005), head of moderate width for genus compared to pronotum (WH/WPm 0.72±0.014), pronotum of moderate width in comparison to elytra (WPm/WE 0.75±0.018).

Color. Body color rufotestaceous, appendages testaceous.

Prothorax. Pronotum slightly long (LP/LE 0.40±0.012) and moderately elongate (WPm/LP 1.25±0.029), with lateral margins rectilinear constricted posteriorly (WPm/WPp 1.56±0.068). Width between anterior angles moderately greater than between posterior angles (WPa/WPp 1.21±0.034).

Elytra. Slightly depressed along suture, comparatively long (LE/SBL 0.52±0.013) and slightly wide (WE/LE 0.67±0.010). Lateral margins slightly divergent at basal third, subparallel at middle and evenly rounded to apex in apical third.

Male genitalia. Median lobe ( Fig. 13View FIGURE 13 A) with straight ventral margin and bent downward enlarged apex with tapered rounded tip. Sclerites of internal sac uneven with long, heavily sclerotized ventral and short, weakly sclerotized dorsal branches, united together basally, shorter than half the distance between apical and basal orifices. Left paramere with long apical constriction ( Fig. 13View FIGURE 13 C). Ring sclerite with transverse handle of moderate length ( Fig. 14View FIGURE 14 J).

Female internal genitalia. Goncoxite 2 with long ensiform setae ( Fig. 15View FIGURE 15 G). Spermatheca sclerotized, strikingly elongate, curved, with two indistinctly split compartments ( Fig. 17View FIGURE 17 A). Spermathecal duct extremely long with numerous coils. Attachments of spermathecal duct and gland to spermatheca moderately distant from each other.

Geographical distribution. This species is known from one locality located in North Otago ( Fig. 18View FIGURE 18 B, yellow triangle).

Way of life. Specimens were collected in the tussockland close to a hilltop.

Relationships. In the hypertrophy of features of male median lobe and spermatheca, members of this species resembles those of Z. lescheni  . However, the ranges of these two species are widely isolated geographically, which suggests only a distant relationships if any. Relationships of Z. carltoni  are not postulated here.


New Zealand Arthropod Collection