Paracymbiomma, Rodrigues & Cizauskas & Rheims, 2018

Rodrigues, Bruno V. B., Cizauskas, Igor & Rheims, Cristina A., 2018, Description of Paracymbiomma gen. nov., a new genus of prodidomid spiders from the Neotropical region (Araneae: Prodidomidae) including a new troglobite species, Zootaxa 4514 (3), pp. 301-331 : 303-304

publication ID

https://doi.org/10.11646/zootaxa.4514.3.1

publication LSID

lsid:zoobank.org:pub:B674EE1E-57C0-4FC5-B855-AED8FA6596EA

DOI

http://doi.org/10.5281/zenodo.5957137

persistent identifier

http://treatment.plazi.org/id/8D1487E6-5419-FFB9-FF2D-F64BFC5F77D0

treatment provided by

Plazi

scientific name

Paracymbiomma
status

gen. nov.

Paracymbiomma   gen. nov.

Etymology. The generic name is a contraction of paracymbium and Lygromma   , since the paracymbium is one of the diagnostic structures of the genus. Gender is neuter.

Type species. Paracymbiomma angelim   (original designation).

Diagnosis. Species of Paracymbiomma   gen.nov. are distinguished from those of other Prodidomidae   genera by the combination of the following characters: male palp with paracymbium and RTA single ( Figs 45 View FIGURES 43–48 , 54 View FIGURES 53–56 , 61 View FIGURES 61–64 ), embolus long and filiform with curved tip ( Figs 43, 47 View FIGURES 43–48 , 49 View FIGURES 49–52 , 61 View FIGURES 61–64 ), conductor sclerotized ( Figs 53, 55 View FIGURES 53–56 , 57) and median apophysis membranous with sclerotized tip ( Figs 53 View FIGURES 53–56 , 49 View FIGURES 49–52 , 61 View FIGURES 61–64 , 69–74 View FIGURES 69–74 ); female epigyne with posteromedian atrium and sclerotized margins ( Figs. 51 View FIGURES 49–52 , 59, 63 View FIGURES 61–64 , 67 View FIGURES 65–68 ), vulva with copulatory ducts long, strongly convoluted laterally with proximal part (closest to copulatory openings) translucent ( Figs 52 View FIGURES 49–52 , 68 View FIGURES 65–68 ); secondary spermathecae elongated ( Figs 60, 64 View FIGURES 61–64 ).

Description. Total length males 1.86–3.9 and females 2.25–4.15. Carapace and legs pale yellow, and abdomen weakly pigmented. Carapace longer than wide, narrower at cephalic region ( Fig. 7 View FIGURES 7–12 ). Fovea present as a narrow dark longitudinal line on posterior half of carapace ( Figs 1–7 View FIGURES 1–6 View FIGURES 7–12 ). Eyes variable in number and size ( Figs 13–18 View FIGURES 13–18 ). Chilum present, entire ( Fig. 8 View FIGURES 7–12 ). Chelicerae relatively long (0.27–0.9) with a convex boss ( Fig. 8 View FIGURES 7–12 ); promargin with 1–4 teeth, retromargin with 3–6 teeth; one large promarginal escort setae at base of fang ( Fig. 31 View FIGURES 31–36 ); comb of whisker setae at cheliceral promargin near the base of fang on all females collected in caves ( Fig. 20 View FIGURES 19–24 ). Endites slightly convergent anteriorly ( Fig. 9 View FIGURES 7–12 ), with dense scopulae on internal margin; serrula present in single row ( Fig. 21 View FIGURES 19–24 ); labium longer than wide ( Fig. 9 View FIGURES 7–12 ). Sternum shield shaped, anteriorly straight, longer than wide, rebordered anteriorly and laterally; posterior region strongly protruding between coxae IV with numerous long and erect setae ( Figs 10 View FIGURES 7–12 , 22 View FIGURES 19–24 , 33 View FIGURES 31–36 ); precoxal triangles present ( Fig. 10 View FIGURES 7–12 ). Pedicel with ventral sclerite triangular. Leg formula 4123. Spination variable. All tarsi scopulate ( Figs 23 View FIGURES 19–24 , 25 View FIGURES 25–30 ). Trichobothria present on dorsal tibia, metatarsi and tarsi, arranged in two rows on tarsi ( Fig. 25 View FIGURES 25–30 ) and one on metatarsi ( Fig. 34 View FIGURES 31–36 ); bothrium with 5–9 deep ridges ( Fig. 26 View FIGURES 25–30 ). Tarsal organ capsulate with oval opening, located dorsally on tarsi. Two pectinate claws; tarsi I–II with long toothed claws, with 6–13 teeth ( Figs. 27 View FIGURES 25–30 , 35 View FIGURES 31–36 ) tarsi III–IV with short toothed claws, with 3–4 teeth ( Fig. 36 View FIGURES 31–36 ). Claw tufts composed of slightly widened tenent setae ( Fig. 27 View FIGURES 25–30 ); presence of classic claw tuft clasper ( Figs 27 View FIGURES 25–30 , 35, 36 View FIGURES 31–36 ) Abdomen oval, longer than wide, covered with sheet-shaped translucent scales ( Figs 29, 30 View FIGURES 25–30 ); sparse tactile hairs with several barbs present; dorsum of abdomen anteriorly with long, dark, posteriorly curved setae; postepigastric invaginations present ( Figs 12 View FIGURES 7–12 , 37 View FIGURES 37–42 ). Six spinnerets ( Figs 11 View FIGURES 7–12 , 38, 39 View FIGURES 37–42 ); ALS elongated (0.6–0.13), separated from each other by one diameter or more ( Fig. 11 View FIGURES 7–12 ), piriform gland spigots elongated with few associated setae ( Fig. 40 View FIGURES 37–42 ), male with one major ampullate gland spigot ( Fig. 40 View FIGURES 37–42 ) and female with two; PLS and PMS conical and short ( Figs. 41, 42 View FIGURES 37–42 ), PLS larger than PMS ( Figs 11 View FIGURES 7–12 , 38 View FIGURES 37–42 ); PLS in male with one minor ampular gland spigot and four aciniform gland spigots ( Fig. 41 View FIGURES 37–42 ), females apparently with one minor ampular gland spigot, five aciniform gland spigots and two cylindrical gland spigots; PMS in male with one minor ampular gland spigot and two aciniform gland spigots, females apparently with one minor ampular gland spigot, four aciniform gland spigots and two cylindrical gland spigots ( Fig. 42 View FIGURES 37–42 ). Female pedipalp with one pectinate claw with 6–8 teeth ( Fig. 28 View FIGURES 25–30 ). Male palp: tibia generally short, except in P. caecus   sp. nov., in which it is more elongated ( Figs 65, 66 View FIGURES 65–68 , 74 View FIGURES 69–74 ), RTA single, directed apically ( Figs 54, 56 View FIGURES 53–56 ); cymbium oval with apical scopula ( Figs 62 View FIGURES 61–64 , 66 View FIGURES 65–68 ), paracymbium present ( Figs 43, 45 View FIGURES 43–48 , 54 View FIGURES 53–56 , 61 View FIGURES 61–64 ); bulb rounded ( Figs 53, 55 View FIGURES 53–56 , 69–72 View FIGURES 69–74 ) or ovoid ( Figs 61 View FIGURES 61–64 , 65 View FIGURES 65–68 , 73, 74 View FIGURES 69–74 ); embolus long with curved tip and wide base, inserted between 9–12 o’clock position ( Figs 55 View FIGURES 53–56 , 71 View FIGURES 69–74 ), or 3 o’clock position ( Figs 61 View FIGURES 61–64 , 74 View FIGURES 69–74 ); conductor sclerotized, with curved tip, inserted

prolaterally ( Figs 47 View FIGURES 43–48 , 55 View FIGURES 53–56 , 70 View FIGURES 69–74 ); median apopohysis inserted distally and with sclerotized tip( Figs 69–74 View FIGURES 69–74 ). Epigyne: posterior margin of epigynal plate anteriorly curved ( Figs 51 View FIGURES 49–52 , 67 View FIGURES 65–68 ) or truncate ( Fig. 59); conspicuous posteromedian atrium with well-defined margins ( Figs 59, 63 View FIGURES 61–64 ). Vulva: secondary spermathecae elongated ( Figs 60, 64 View FIGURES 61–64 ); fertilization ducts antero-lateral.

Distribution. Brazil: states of Pará (exclusively in ferruginous caves), Paraíba, Pernambuco, Bahia and São Paulo ( Fig. 75 View FIGURES 75 ).

Composition. Six species: Paracymbiomma angelim   sp. nov.; P. doisirmaos   sp. nov.; P. pauferrense   sp. nov.; P. carajas   sp. nov.; P. bocaina   sp. nov.; P. caecus   sp. nov. ( Figs 1–6 View FIGURES 1–6 ).