Dasymutillini Brothers & Lelej

Brothers, Denis J. & Lelej, Arkady S., 2017, Phylogeny and higher classification of Mutillidae (Hymenoptera) based on morphological reanalyses, Journal of Hymenoptera Research 60, pp. 1-97: 11

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Dasymutillini Brothers & Lelej

trib. n.

Dasymutillini Brothers & Lelej   trib. n.

Type genus.

Dasymutilla   Ashmead, 1899. This group is paraphyletic in most analyses, although, interestingly, monophyletic in the tree derived from males only (Fig. 10) and that from the matrix with duplicated terminals reflecting maximum polymorphisms (Fig. 7), and almost so in the tree derived from the reduced matrix in which the most polymorphic characters had been deleted (Fig. 8). Using Fig. 13 as the base, moving the terminals to reflect the arrangement in the preferred tree (Fig. 5) (except in retaining the Euspinolia   - Hoplocrates   group as sister to the remaining pseudomethocines) actually added four steps, making the proposed final arrangement preferable in this regard. The group is not supported by resampling nor by any unique and unambiguously placed synapomorphies, but there is a single unique but ambiguously placed synapomorphy for both additive and non-additive characters: 10.2, eye strongly convex in females (but also in several other sphaeropthalmines and Seyrigilla   , and less convex in Odontomyrme   ). There are also some ambiguously placed homoplasious synapomorphies, the most significant being: 135.2, mesoscutal notaulus absent in winged males (but also in most pseudomethocines, a few sphaeropthalmines s.s. and scattered terminals elsewhere, and present in Gogoltilla   and Tobantilla   ). It is not surprising that Dasymutilla   was shown to be paraphyletic in the analysis of duplicated terminals (Fig. 7), since it is generally recognized that the genus is highly variable (and even very difficult to separate from Traumatomutilla   Andr√©), although recent reviews have not suggested the recognition of further genera or even subgenera; we tried to capture some of that variability in the selection of exemplars. The tribe is Neotropical, Australian and Nearctic in distribution, with 24 sub/genera; females are known for 100% and males for 95% of those taxa.