Pterostichus (Nialoe) chokaisanus Sasakawa, 2009

Sasakawa, Koji & Mitsuduka, Yoshiji, 2023, A morphology-based revision and phylogenetic analysis of the Pterostichus macrogenys species group (Coleoptera, Carabidae) and implications for differentiation of the species group, Deutsche Entomologische Zeitschrift 70 (2), pp. 291-310 : 291

Pterostichus (Nialoe) chokaisanus Sasakawa, 2009

Figs 3A, B , 5A-H , 6I, J

Materials examined.

1♂ 2♀, Momoyake , alt. 1030 m, Yurihonjo-shi, Akita Prefecture, Japan (MGRS: 54 SVJ26680 29227; 39.1091115°N, 140.15197046°E), 24. vi-15. vii. 2021, subterranean baited trap, Y. Mitsuduka leg. (Figs 3A , 5A-D , 6I ) ; 1♂, Zaô-chûô-kôgen, Zao-onsen Spa , Yamagata-shi, Yamagata Prefecture, Japan (MGRS: 54 SVH49227 23449; 38.15750459°N, 140.42046179°E), 1-6. ix. 2020, subterranean baited trap, Y. Mitsuduka leg. (Figs 3B , 5E-H , 6J ) .

Notes.

The endophallus of the Zaô-chûô-kôgen specimen could not be inflated sufficiently due to changes in the membranous part (Fig. 5E-H ), but the specimen was identified unambiguously as P. chokaisanus , based on species-specific structures of the endophallus. The body lengths (mm) of the Momoyake specimens (♂/♀/♀) are BLm 14.91/14.24/14.94, BLl 13.65/13.08/13.68 and BLc 13.18/12.56/13.21; and those of the Zaô specimen are BLm 14.20, BLl 12.83 and BLc 12.40. The Zaô-chûô-kôgen specimen is the southernmost distribution record of this species.

Figures

Figure 3. Habitus dorsal view of the Pterostichus macrogenys species group. A. Pterostichus chokaisanus male from Momoyake; B. P. chokaisanus male from Zao-chuo-kogen; C. P. kitakamisanus male from Odateminami; D. P. kitakamisanus holotype male from Genbeidaira.

Figure 5. Endophallus of Pterostichus chokaisanus male from Momoyake (A-D), P. chokaisanus male from Zao-chuo-kogen (E-H), P. kitakamisanus male from Odateminami (I-L) and P. kitakamisanus holotype male from Genbeidaira (M, N). Fully inflated endophallus in left lateral (A, E, I, M), apical (B, F, J, N), right lateral (C, G, K) and basal part in dorsal and apical part in ventral (D, H, L) views.

Figure 6. Right paramere of Pterostichus namahage sp. nov. holotype male from Haraikawa (A), P. kamurosanus sp. nov. holotype male from Ariya (B), P. atsumidakensis sp. nov. holotype male from Take-no-koshi (C-E), P. asahinus male from Momoyake (F-H), P. chokaisanus male from Momoyake (I), P. chokaisanus male from Zao-chuo-kogen (J), P. kitakamisanus male from Odateminami (K, L) and P. kitakamisanus holotype male from Genbeidaira (M). Left lateral (A-C, F, I-K), right lateral (L, M), apical part in apical and basal part in ventral (D, G) and apical part in dorsal (E, H) views.