Antikuna cimrmani Kaderka, Ferretti & Hüsser, 2021

Kaderka, Radan, Ferretti, Nelson, Hüsser, Martin, Lüddecke, Tim & West, Rick, 2021, Antikuna, a new genus with seven new species from Peru (Araneae: Theraphosidae: Theraphosinae) and the highest altitude record for the family, Journal of Natural History 55 (21 - 22), pp. 1335-1402 : 1384-1392

Antikuna cimrmani Kaderka, Ferretti & Hüsser , sp. nov.

( Figures 43–45 and 49 , Table 11)

Types: Female holotype (MUSM-ENT 0514017, formerly RKCP 0779) from Peru,

Huancavelica, Colcabamba, 3,980 m a.s.l., 2011, M. Černička col.

Etymology: The specific name is dedicated to Jára Cimrman, one of the greatest Czech playwrights, poets, composers, teachers, travellers, philosophers, inventors, detectives, mathematicians, and sportsmen of the 19th and early 20th century.

Diagnosis: Females of Antikuna cimrmani sp. nov. can be distinguished from all other congeners, except for A. urayrumi sp. nov. and A. valladaresi sp. nov., by dark grey legs with darker femora. Females also differ from females of A. urayrumi sp. nov. and A. valladaresi sp. nov. in blue iridescence of femora. Females also differ from A. cernickai sp. nov. in the shape of seminal receptacles (long and narrow in Antikuna cimrmani sp. nov. and short and wide in A. cernickai sp. nov.).

Distribution ( Figure 49 ): Known only from the type locality, only 14 km distant from the type locality of Antikuna cernickai sp. nov.

FEMALE (MUSM-ENT 0514017, formerly RKCP 0779) ( Figures 43–45 ): Total length: 23.1, carapace length 11.1, width 10.1, caput domed, chelicerae with 12 teeth on promargin. Cheliceral teeth pattern from basal end: right side: V-V-vvvvvvVV-V-V, 6 big and 6 small teeth. Left side: v-V-V-vvvvVVVV-V, 7 big and 5 small teeth. Anterior eye row procurved, posterior eye row recurved. Eye sizes and interdistances ( Figure 44 (c)): AME 0.27 (circular), ALE 0.36 (oval), PME 0.38 (drop-shaped), PLE 0.38 (oval), AME-AME 0.33, AME-ALE 0.17, PME-PME 0.75, PME-PLE 0.13, ALE-PLE 0.32, AME-PME 0.18, OQ length 0.92, width 1.66. Ocular tubercle: length 1.38, width 1.66, clypeus narrow, length 0.182. Fovea transverse, procurved, width 2.47, 7.64 from anterior edge of carapace. Labium length 1.5, width 2.3, anterior third with 13 cuspules, maxillae with 137 – 156 cuspules in basal half and short spiniform setae in apical half. Sternum domed, length 5.8, width 4.9, with three visible pairs of sternal sigilla located near coxae III (length 0.44, 0.55 from edge of sternum), coxae II (length 0.29, 0.37 from edge of sternum) and coxae I (length 0.25, 0.16 from edge of sternum). Leg pattern: IV>I>II>III, legs I and IV almost equal. All leg segments uniform.

Scopulae: All tarsi 100% densely scopulate, metatarsi I 100%, metatarsi II 70%, metatarsi III 50%, metatarsi IV 30% scopulate. Tarsal scopulae I integral, tarsal scopulae II divided by a longitudinal band of setae, in tarsi III and IV divided by a wide band of setae. Dorsal face of all tarsi and cymbium with two irregular longitudinal rows of very short claviform trichobothria. Denticulation pattern of paired tarsal claws on left legs (prolateral/retrolateral row): I 5/0, II 5/5, III 5/4, IV 3/4. Plumose setae on retrolateral face of femur IV absent.

Spination: femora I p 0-0-1, II p 0-1-1, III p 0-1-1, d 0-0-1, IV d 0-0-1 and femora of palps p 0-0-1, patellae I – IV and patellae of palps 0, tibiae I v 0-0-1 (apical), II v 0-1-3 (apical), p 1-1-0, III v 1-1-2 (apical), p 1-1-0, r 1-0-1, IV v 1-1-3 (apical), r 1-0-1 and tibiae of palps v 1-1-2 (apical), metatarsi I v 0-0-3 (apical), II v 3-0-4 (apical), III v 1-3-10 (apical), p 0-1-1, r 0-1-1, IV v 2-3-3-2-7 (apical), p 0-0-1, r 0-1-1, tarsi I – IV and tarsi of palps 0.

Spermathecae ( Figure 45 ): two separated seminal receptacles with subapical constriction.

Abdomen ( Figure 44 (d, e)): urticating setae of type III are located in oval central patch . PLS: length 4.83, basal segment 2.08, middle segment 1.22, apical segment 1.53, all digitiform. PMS: 1.27.

Colouration and covering setae: dorsal view ( Figure 43 ): carapace grey, caput dark grey, coxae and trochantera grey, chelicerae grey, femora dark grey with blue iridescence, patellae, tibiae,metatarsi and tarsi grey, intermixed with long, pale setae.Patellae I, II, palpal patella and tibiae with two almost equal parallel longitudinal stripes without covering setae, patellae III, IV with two unequal diagonal stripes. Palpal femur prolaterally partly bare. Abdomen covered with short dark grey setae, intermixed with long, reddish-brown setae, except oval central patch which is brown and covered with long reddish-brown setae, arranged diagonally to the longitudinal body axis. Length of central patch: 6.5, width 7.3. Ventral view: labium and maxillae reddish-brown, sternum, coxae, trochantera, femora, patellae, tibiae and metatarsi grey. Abdomen ventrally without dark longitudinal band. Spinnerets light brown.

MALE: unknown.

Figures

Figure 43. Antikuna cimrmani sp. nov. from Peru, Huancavelica, Colcabamba, 3,980 m a.s.l., habitus of female holotype (MUSM-ENT 0514017, formerly RKCP 0779), dorsal view. Scale bar = 10 mm. Photo by R. Kaderka.

Figure 44. Antikuna cimrmani sp. nov., female holotype (MUSM-ENT 0514017, formerly RKCP 0779) from Peru, Huancavelica, Colcabamba, 3,980 m a.s.l. (a). Carapace, coxae and trochantera, dorsal view. (b). Sternum, labium, maxillae and coxae, ventral view. (c). Ocular tubercle, dorsal view. (d). Abdomen, dorsal view. (e). Abdomen, ventral view. (f). Left maxilla with short spiniform setae in apical half, ventral view. Scale bar = 1 mm (a, b, c, f). Scale bar = 10 mm (d, e). Photo by R. Kaderka.

Figure 45. (a, c). Antikuna cimrmani sp. nov., adult female holotype (MUSM-ENT 0514017, formely RKCP 0779) from Peru, Huancavelica, Colcabamba, 3,980 m a.s.l. Spermathecae with two separated, basally enlarged seminal receptacles. Spermathecae dissected from specimen in alcohol. (a). Ventral view. (b). Dorsal view. (c). Spermathecae dissected from exuvia, ventral view. Scale bar = 1 mm. Photo by R. Kaderka.

Figure 49. Distribution map of Antikuna gen. nov. in Peru. Prepared by M. Hüsser.