Furcadia Opitz, 2017

Opitz, Weston, 2017, Classification, Natural History, and Evolution of the Orthopleurinae (Coleoptera: Cleridae). Part VI. The New Genera Coadnatus Opitz, Furcadia Opitz, and Latupusillus Opitz, One New Species of Funicula Opitz, and Four New Species of Romanaeclerus Winkler, The Coleopterists Bulletin 71 (3), pp. 421-433 : 426-428

Furcadia Opitz , new genus

Type Species. Furcadia sandala Opitz , new species.

Diagnosis. In a key to the genera of Orthopleurinae ( Opitz 2009a) , Furcadia keys to the Old World genus Funicula , from which Furcadia is distinguished by a more robustly developed antennal capitulum (compare Figs. 3 and 8 ), subsecuriform maxillary and labial terminal palpomeres, and a well-diffentiated pronotal arch (compare Figs. 21 and 23 ). Also, the spicular apodemes are not contiguous in Furcadia ( Fig. 13 ), whereas in Funicula these apodemes are contiguous for nearly their entire length ( Fig. 12 ).

Synapotypic Characteristic. Intraphallic-plate membrane spinose.

Description. Size: Length 4.8–8.0 mm; width 1.2–2.0 mm. Form: Oblong-rectangulate, about 3.5 times longer than broad. Vestiture: Discs of cranium and pronotum vested with pale setae, elytral disc vested with 1° setae and shorter, profusely distributed 2° setae. Head: Cranium quadrate, frons wide, indented with small, contiguous, setiferous punctation; gula small, triangular, sutures oblique, gular process narrow and bifid; labrum short, medial incision deep; mandible short, subacuminate; maxillary and labial terminal palpomeres subsecuriform; eyes small, finely faceted, ocular notch large; antenna ( Figs. 3, 5 ) capitate, capitulum lax, scape elongate, longer than combined length of pedicel and antennomere 3, funicular antennomeres progressively shorter to capitulum, capitular antennomeres 9 and 10 sexually dimorphic, triangular in females, oblong-triangular in males, antennomere 11 oblong. Thorax: Pronotum ( Fig. 23 ) quadrate, with tumescences on pronotal disc, lateral margins slightly incised at anterior third, arch welldeveloped, remainder of pronotum finely punctate, transverse subapical and prebasal depressions welldeveloped, prointercoxal process expanded distally; pronotal projections long, contacting but not fused with prointercoxal process; elytron sculptured with asetiferous punctation that diminish after basal half, punctuation subseriate in basal half, 1° setae always adjacent to asetiferous punctation, 2° setae present, epipleural fold narrowing to elytral apex, anterior margin carinate; tibial spur formula 2-2-2, tarsal pulvillar formula 3-3-3, unguis with well-developed

36) Romanaeclerus vitellinus .

denticle. Abdomen: Six visible sternites; aedeagus ( Figs. 14, 15 ) with lateral lobes feebly developed; phallobasic struts not confluent with phallobasic apodeme, phallobasic rod broad; phallic plates narrow; spicular plates flared ( Fig. 13 ), plates slightly truncate, spicular apodemes not fused, intraspicular plate oblong-linear.

Distribution. This monotypic genus is known from Laos.

Etymology. The genus name Furcadia is a compound name that stems from the Latin furca (= fork) and the Greek dia (= between). I refer to the complete separation of the spicular apodemes. Gender: Feminine.

Furcadia sandala Opitz , new species ( Figs. 3, 5, 13–15 , 23 , 31 , 39 )

Type Material. Holotype: ♀. Phou Pan (Mt.), Houa Pan Prov., NE LAOS, 10–22.V.2011, St Jakl leg ( FSCA) . Paratypes: Two specimens. LAOS, Phongsaly Province, 21°41 ′ N 102°6E ′, Phongsaly environs, 6–17-V-2004, 1500 m, M. Brancucci ( RGCM, 1; WOPC. 1) .

Diagnosis. This is the only species in the genus, so it is recognized by the combination of characteristics defining the genus.

Description. Size: Length 4.8 mm; width 1.2 mm. Form: As in Fig. 31 . Color: Black, except frons, mouthparts, prosternum, pterothoracic venter, and abdomen dark castaneous; pronotal arch black, pronotal disc red. Head: Antennal capitulum welldeveloped ( Figs. 3, 5 ), capitular antennomeres 9 and 10 triangular, antennomere 11 oblong-rectangular, funicular antennomeres subfiliform; maxillary and labial terminal palpomeres digitiform; eyes much narrower than frons (EW/FW = 15/30). Thorax: Pronotum ( Fig. 23 ) quadrate (PW/PL = 70/70), lateral margins incised at apical 1/3; elytral asetiferous punctation small and shallow, extending to just beyond elytral middle, interstitial spaces arenose, epipleural fold narrowing to elytral apex. Abdomen: Pygidium scutiform; aedeagus ( Figs. 14, 15 ) with lateral lobes feebly developed; phallobasic struts not confluent with phallobasic apodeme, phallobasic rod broad; phallic plates narrow; spicular plates flared ( Fig. 13 ), plates slightly truncate, spicular apodemes not fused, intraspicular plate oblonglinear.

Natural History. The type specimens were collected during May, two at 1,500 m.

Variation. Size: Length 4.8–8.0 mm; width 1.2–2.0 mm. Except for body size, the type specimens are quite homogeneous.

Distribution. This species is known only from Laos ( Fig. 39 ).

Etymology. The Latin specific epithet sandala (= slipper or sandal) refers to the body form of this beetle.

Figures

Figs. 1–20. Antennae and male genitalia. Antennae: 1) Coadnatus sakaii; 2) Romanaeclerus gerstmeieri; 3) Furcadia sandala, female; 4) Romanaeclerus quadratus; 5) F. sandala, male; 6) Romanaeclerus dimidius; 7); Romanaeclerus vitellinus; 8) Funicula albocapilla; 9) Latupusillus laculatus. Aedeagi and spicular fork: 10) F. albocapilla, phallus; 11) F. albocapilla, tegmen; 12) F. albocapilla, spicular fork; 13) F. sandala, spicular fork; 14) F. sandala, phallus; 15) F. sandala, tegmen; 16) C. sakaii, tegmen, dorsal view; 17) C. sakaii, phallus; 18) C. sakaii, tegmen, ventral view; 19) R. vitellinus, phallus; 20) R. vitellinus, tegmen, ventral view.

Figs. 21–28. Pronota, dorsal view. 21) Funicula albocapilla; 22) Romanaeclerus quadratus; 23) Furcadia sandala; 24) Romanaeclerus vitellinus; 25) Coadnatus sakaii; 26) Latupusillus laculatus; 27) Romanaeclerus dimidius; 28) Romanaeclerus gerstmeieri.

Figs. 29–32. Habitus. 29) Coadnatus sakaii; 30) Funicula albocapilla; 31) Furcadia sandala; 32) Latupusillus laculatus.

Figs. 39–40. Geographic distributions of 39) Coadnatus sakaii and Furcadia sandala; 40) Funicula albocapilla.