Verodes inaequalis (Say)

Foley, Ian A. & Ivie, Michael A., 2008, A phylogenetic analysis of the tribe Zopherini with a review of the species and generic classification (Coleoptera: Zopheridae), Zootaxa 1928 (1), pp. 1-72 : 54-55

Verodes inaequalis (Say)

Figs. 56–57

Eurychora inaequale Say, 1835: 182 .

Nosoderma inaequale ; Gebien, 1936: 669. García-París et al., 2006: 230.

Nosoderma inaequalis ; Casey, 1907b: 470. Blackwelder, 1945: 514.

Nosoderma morbillosum Solier, 1841: 37 . (Synonymy by Gebien, 1936: 669.)

Nosoderma morbillosa ; Casey, 1907b: 475. Ṡ lipi ń ski and Lawrence, 1999: 13.

Nosoderma vicinum Solier, 1841: 38 . (Synonymy by Gebien, 1936: 669.)

Nosoderma vicina ; Casey, 1907b: 476.

Nosoderma prominens Casey, 1907b: 478 . Gebien, 1936: 669. Blackwelder, 1945: 514. García-París et al., 2006: 230. NEW SYNONYMY.

Verodes prominens ; Foley and Ivie, 2007: 70.

Nosoderma senex Casey, 1907b: 479 . Gebien, 1936: 669. Blackwelder, 1945: 514. García París et al., 2006: 230. NEW SYNONYMY.

Verodes senex ; Foley and Ivie, 2007: 70.

Nosoderma brevicollis Casey 1907b: 478 . Blackwelder 1945: 514. García-París et al., 2006: 229. NEW SYNONYMY.

Nosoderma brevicolle ; Gebien, 1936: 669.

Verodes brevicollis ; Foley and Ivie, 2007: 71.

Nosoderma subglabra Casey, 1907b: 478 . Blackwelder, 1945: 514. NEW SYNONYMY.

Nosoderma subglabrum ; Gebien, 1936: 669. García-París et al., 2006: 230.

Verodes subglabrus ; Foley and Ivie 2007: 70 .

Diagnosis: This species can be distinguished by the ridge in the 5 th elytral interval not reaching the base of the elytra, and strongly expanded hind angles of the pronotum lacking the very large conical nodules seen in V. scabrosus .

Distribution: Central Mexico north through the Sierra Madre Occidental at elevations between 2,000 – 4,000 meters.

Recorded distribution: A summary of the 284 specimens examined is MEXICO: DISTRITO FED- ERAL, DURANGO, GUANAJUATO, GUERRERO, HIDALGO, JALISCO , MÉXICO, MORELOS , PUE- BLA, SINALOA, SONORA, VERACRUZ .

Types: Eurychora inaequale: The confusion surrounding the species epithet inaequale of Dejean and inaequalis of Say and the fact that Say’s types have been lost requires the designation of a Neotype for Eurychora inaequalis Say 1835 . Casey (1907b) recognized this need and identified a specimen which is here designated the NEOTYPE of Eurychora inaequalis . NEOTYPE: ♀, intact in NMNH. Mex./ Casey; bequest; 1925/ Eurychora ; inaequalis; Say/ NEOTYPE; Eurychora ; inaequalis ; ♀ Say, 1835; desg. Foley & Ivie ’07.

Nosoderma prominens : LECTOTYPE, here designated: ♀, missing right mesotarsus, in NMNH. Omilteme,; Guerrero,; 8000 ft.; H. H. Smi/ Casey; bequest; 1925/ N. inaequale Say ; morbillosum/ Type NMNH; 46386/ prominens; Csy./ red rectangle label Lectotype; Nosoderma ; prominens ; ♀ Casey, 1907; desg. Foley & Ivie ’07.

Nosoderma senex : LECTOTYPE, here designated: ♂, missing left meta leg, in NMNH. Las Vigas; Mexico; Hoege/ Casey; bequest; 1925/ Type NMNH; 46384/ senex; Csy./ red rectangle label Lectotype; Nosoderma ; senex ; Casey, 1907; desg. Foley & Ivie ’07.

Nosoderma brevicollis : LECTOTYPE, here designated: ♂, intact, in NMNH. Mex./ Casey; bequest; 1925/ Type NMNH; 46387/ brevicollis; Csy./ red rectangle label Lectotype; Nosoderma ; brevicollis ; ♂ Casey, 1907; desg. Foley & Ivie ’07.

Nosoderma brevicollis : LECTOTYPE, here designated: ♂, missing right antenna and protarsus, in NMNH. Guer./ Casey; bequest; 1925/ Type NMNH; 46385/ Nosoderma ; subglabra; Csy./ red rectangle label Lectotype; Nosoderma ; subglabra ; ♂ Casey, 1907; desg. Foley & Ivie ’07. PARALECTOTYPE, here designated: ♂, intact in NMNH. Same data as Lectotype.

Notes: One of the most commonly collected, widespread, and morphologically diverse members of the genus. Some of this variation is represented in the Casey types; this includes variation in the expansion of the hind angles of the pronotum, strength of the elytral punctures, and general shape of the elytral apex. All of the variation seen within the Casey types is well within the range for the species throughout its distribution.

Figures

FIGURES 54–56. 54–55. Hypomera. 54. Sesaspis sylvatica; 55. Verodes exsculptus. 56. Ventral surface of head and pronotum of Verodes inaequalis. Scale bars = 1mm.

FIGURES 57–60. Dorsal habiti. 57. Verodes inaequalis, Mexico, Puebla.; 58. V. exsculptus, Mexico, Oaxaca; 59. V. asperatus, Mexico, Hidalgo; 60. V. scabrosus, Mexico. Scale bars = 5mm.