Vespiodes phaios, Dikow, Torsten, 2010
treatment provided by
phaios Greek adjective = brown. Referring the overall brown coloration of this attractive species.
The species is distinguished from congeners by the predominantly apruinose and asetose scutum, the asetose abdominal tergites, features of the male terminalia, and its apparent distribution in south-eastern coastal Kenya.
Head: brown, in general silver pruinose; width distinctly greater than thorax, interocular distance on vertex larger than at ventral eye margin, vertex between compound eyes slightly depressed, parafacial area less than ½ the width of central facial gibbosity; facial gibbosity distinct, well-developed and discernible in lateral view; mystax white, covering only lateral facial gibbosity (asetose medially); frons predominantly apruinose, vertex apruinose, postgenae lightly silver pruinose; setation: vertex white, frons white, ocp setae brown or white, pocl setae white; ocellar triangle apruinose; proboscis brown, long, projecting beyond fronto-clypeal suture; labella large, much wider than prementum, about ½ length of prementum, unsclerotised laterally; maxillary palpi laterally compressed (leaflike), brown or light brown, shorter than length of pedicel.
Antenna: brown or orange, scape and pedicel white setose dorsally and ventrally; postpedicel cylindrical in proximal ½, symmetrically bulbous in distal ½, ≥ 11.0 times as long as combined length of scape and pedicel; apical ‚seta-like‘ sensory element situated apically in cavity on postpedicel.
Thorax: brown, predominantly apruinose; scutum medially dark brown, laterally brown, surface macrorugose, predominantly apruinose, small antero-lateral spots and extreme lateral margins (dorsal to supero-posterior anepisternum) silver pruinose, scutal setation comprised of distinct rows of long dorsocentral setae and lateral scutal setae; dc setae pre- and postsuturally white, acr setae absent, lateral scutal setae white, npl, spal, and pal setae absent; postpronotal lobe light brown, partly silver pruinose; proepisternum, lateral postpronotum, and postpronotal lobes long brown setose or long white setose; scutellum apruinose, asetose, apical scutellar setae absent; mesopostnotum, anatergite, and katatergite apruinose, mesopostnotum asetose, anatergite asetose, katatergite long white setose; katatergite ± flat; anterior anepisternum asetose, supero-posterior anepisternum asetose; posterior anepimeron asetose, katepimeron asetose; metepimeron evenly elevated, same colour as T1, grey pruinose, asetose; metepisternum grey pruinose, asetose.
Leg: brown, setation predominantly brown; pro coxa apruinose, short white setose, mes coxa apruinose, short white setose, met coxa grey pruinose, white setose; met trochanter setose medially; femora brown, met femur proximally yellow, met femora distinctly clubbed in distal ⅔, in distal ½ macrosetose, 1 antero-ventral and 1 postero-ventral row of macrosetae; all tibiae laterally arched, met tibia cylindrical, ventral keel absent; pro and mes tarsomere 1 as long as combined length of tarsomeres 2-3, met tarsomere 1 slightly longer than tarsomere 2, tarsomeres 1 and 2 longer than tarsomeres 3 and 4 combined; pulvilli well-developed, as long as well-developed claws, and as wide as base of claws; empodium absent.
Wing: length = 15.9 mm; slightly brown stained, darker brown stained anterior to and distal to M veins, veins brown, microtrichia absent; cells r1, r4, r5, m3, + cup closed; C terminates at junction with R1; R4 terminates in R1; R5 terminates in R1; stump vein ( R 3) at base of R4 present, long but not reaching R2; R4 and R5 widest apart medially; r-m distinct, R4+5 and M1 apart, connected by crossvein; M1 curves anteriorly at r-m, M1 (or M 1+ M 2) terminates in R1; CuA1 and CuA2 split proximally to m-cu (cell m3 narrow proximally); M3+CuA1 do not terminate together in C; A1 undulating, cell a1 wide, A1 and wing margin further apart proximally than distally, alula well-developed; halter brown.
Abdomen: brown and yellow; setation comprised of scattered white and brown setae, surface entirely smooth; T1 brown, T2-4 brown with yellow anterior margins; T1-3 sparsely white setose; T predominantly apruinose; S1 brown, S2-4 brown with yellow anterior margins; S1 asetose, S2-3 sparsely white setose; S predominantly apruinose; T2-4 distinctly narrowed, forming a waist, abdomen vespiform; bullae on T2 brown, transversely elongate, surface entirely smooth, T2 surface anterior to bullae smooth.
Male terminalia: T1-7 well-developed, entirely sclerotised, T8 postero-medially weakly sclerotised, with anterior transverse sclerotised bridge connecting lateral sclerites; T7-8 anteriorly with 2 lateral apodemes; S6 regular, without any special setation postero-medially, S8 well-developed and simple, not fused to T8 dorso-laterally, entire (undivided) ventro-medially; epandrium formed by single sclerite (fused medially ± entirely), blunt, evenly rounded; subepandrial sclerite without lateral or median protuberances; hypandrium strongly concave, cup-shaped, entirely sclerotised ventrally (forming a single sclerite), partially fused with gonocoxite anteriorly; gonocoxites dorso-ventrally flattened in distal ½, higher in proximal ½, without median or lateral protuberance, gonocoxal apodeme present, short (at most slightly extending hypopygium anteriorly); 2 functional aedeagal prongs, short and wide, medio-distally connected; aedeagal epimere present, distally simple, evenly rounded; lateral ejaculatory processes absent; ejaculatory apodeme formed by single dorso-ventrally oriented plate; ventro-median margin of dorsal aedeagal sheath heavily sclerotised (appearing entirely closed); dorsal aedeagal sheath long, sperm sac entirely covered; sperm sac appearing ± heavily sclerotised.
Type locality and distribution:
Muhaka Forest (04°19'29" S; 039°31'27" E), Kenya (Fig. 48). Biodiversity hotspot/high-biodiversity wilderness area: Coastal Forest of Eastern Africa/-.
Kenya, Nairobi, National Museum of Kenya
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