Onthophagus bicolor Raffray, 1877

Ziani, Stefano, Abdel-Dayem, Mahmoud Saleh, Aldhafer, Hathal Mohammed & Barbero, Enrico, 2019, An overview of the Onthophagini from the Arabian Peninsula (Coleoptera: Scarabaeoidea: Scarabaeidae), Zootaxa 4658 (1), pp. 1-36 : 23

Onthophagus bicolor Raffray, 1877

[ Fig. 19 ]

Literature quotes. YEMEN: Aden governorate, Aden. El-Hadj [locality not found] (d’Orbigny 1913a). Without locality (Scott in Paulian 1948) .

Material examined. YEMEN: “ El-Hady ” [without further data, locality not found], C. Martin leg. 2 exx. ( NHMUK) ; Arabia / Yemen [without further data], J. G. Millingen leg. 6 exx. ( NHMUK) . OMAN: Dhofar Governorate, Ain Razat , 110 m (1707′75N 5414′24E), 21.ix.2001, M. Dellacasa leg. 1 ex ( MSNC) ; Al Mughsayl , env., 42 m (1653′74N 5347′90E), 28.viii.2000, M. Dellacasa leg. 3 exx. ( MSNC) ; ibidem (1653′02N 5347′90E), 5.ix.2000, M. Dellacasa leg. 1 ex. ( MSNC) ; ibidem, 8.IX.2000, M. Dellacasa leg. 5 exx. ( MSNC) ; ibidem, 11.ix.2000, M. Dellacasa leg. 24 exx. ( MSNC) ; ibidem, 40 m (1653′83N 5346′31E), 6.ix.2002, M. Dellacasa leg. 6 exx. ( MSNC) ; ibidem, 9.ix.2002, M. Dellacasa leg. 3 exx. ( MSNC) ; ibidem, 12.ix.2002, M. Dellacasa leg. 3 exx. ( MSNC) ; Khor Rori , env., 36 m (1702′27N 5426′16E), 24.viii.2000, M. Dellacasa leg. 7 exx. ( MSNC) ; Rakhyut , env. (1645′00N 5324′67E), 12.ix.2002, M. Dellacasa leg. 7 exx. ( MSNC) ; road Salalah-Darbat , 23 m (1703′14N 5425′58E), 9.ix.2000, M. Dellacasa leg. 5 exx. ( MSNC) ; road 41, Al Mughsayl , env. 30 m (1653′71N 5349′01E), 23.ix.2001, M. Dellacasa leg 1 ex. ( MSNC) ; ibidem, 5.x.2001, M. Dellacasa leg. 1 ex. ( MSNC) ; road 41, between Raysut and Adwnab , 90 m (1657′72N 5352′73E), 23.ix.2001, M. Dellacasa leg. 6 exx. ( MSNC) ; road 47, Adwnab , env., 113 m (1655′34N 5350′56E), 5.ix.2000, M. Dellacasa leg. 18 exx. ( MSNC) ; road 47, ante Al Mughsayl , 60 m (1654′11N 5349′20E), 11.ix.2000, M. Dellacasa leg. 52 exx. ( MSNC) ; ibidem, 11.ix.2000, D. Gianasso leg. 15 exx. ( MSNC) ; Wadi Ashawq , 40 m (1653′83″N 5346′31″E), 23.ix.2001, M. Dellacasa leg. 3 exx. ( SZCM) ; Wadi Ashawq , 57 m (1653′88N 5346′31E), 5.ix.2000, M. Dellacasa leg. 5 exx. ( MSNC) ; ibidem, 11.ix.2000, M. Dellacasa leg. 54 exx. ( MSNC) ; ibidem, 40 m (1653′83N 5346′31E), 23.ix.2001, M. Dellacasa leg. 66 exx. ( MSNC, SZCM) ; Wadi Darbat (1704′50N 5425′88E), 9.ix.2000, M. Dellacasa leg. 2 exx. ( MSNC) .

Distribution. Arabian Peninsula: Yemen (d’Orbigny 1913a). Africa: Eritrea, South Africa, Tanzania (d’Orbigny 1900), Somalia (d’Orbigny 1913a), Ethiopia, Nigeria, Sudan ( Arrow 1931), Asia: Pakistan ( Ziani & Bezděk 2016).

Remarks. First record for Oman. All the literature records for Saudi Arabia ( Arrow 1931; Scott in Paulian 1948; Baltahsar 1963; Zunino 1975, 1981) are without precise locality, and could be due to a wrong reading of d’Orbigny (1913a) who reported: “ Arabia : Yemen, Aden ”. Therefore, we herein choose to exclude this species from the fauna of the Saudi Arabia , until proven otherwise.

Figures

FIGURES 19–26. Onthophagini of Arabian Peninsula. 19, Onthophagus bicolor Raffray, 1877: OM (Dhofar) Wadi Ashawq; 20, Onthophagus latipennis d’Orbigny, 1898: OM (Dhofar) Al Mughsayl; 21, Onthophagus trapezicornis d’Orbigny, 1902: SA (Bahah) 5 km N Al Makhwah; 22, Onthophagus brittoni Paulian, 1948, ♂: SA (Asir) Abha, Raydah; 23, Onthophagus brittoni Paulian, 1948, ♀: SA (Asir) Abha, Raydah; 24, Onthophagus brittoni Paulian, 1948, paramera, lateral view, 25, Onthophagus brittoni Paulian, 1948, parameres, dorsal view; 26, Onthophagus brittoni Paulian, 1948, internal sac: accessory lamellae (above) and lamina copulatrix (below), ventral view. Photos by Gabriele Fiumi.