Hexachaetus mulan, Tian, Mingyi & Deuve, Thierry, 2016

Tian, Mingyi & Deuve, Thierry, 2016, Re-definition and review of the Oriental genus Hexachaetus Chaudoir, 1871 (Coleoptera: Carabidae: Orthogoniini), Zootaxa 4169 (3), pp. 540-554 : 553

Hexachaetus mulan n. sp.

( Figure 11 )

Material. Holotype male, “ Malaysia W., Perak, 30 km SE of Ipoh, 900 m, Cameron Highland, Ringlet , 26- 31.iii.2000, P. Cechovsky leg.”, in CAD, Prague ; paratypes: 1 female, “ Malaysia W., Kelantan Road between Kampong Raja & Gua Musang , 1400–1700 m (Landang Pandrak), 1–28.iv.2006, Cechovsky Petr leg.”, and with an identify label “ Hexachaetus sp. n.? det. Dr. A. Dostel, 2012” ; 1 female, “ Malaysia W., Pahang 30 km SE of Ipoh, 1500 m, Banjaran Titi Wangsa, Tanah Rata , 14–15.iii. 2002, P. Cechovsky leg.”, in CAD and MNHN respectively.

Diagnosis. Head dark brown, pronotum and elytra pale yellow; frontal impressions wide and very deep, strongly sinuate, ending before posterior margin of eyes, frons deeply furrowed medially, as long as frontal impressions, but nearly straight; clypeus deeply concave medially; elytra with apical margin not obliquely truncate, outer angle round, inner angles obtuse though almost rectangular.

Length: 16.5–18.0 mm; width: 6.5–7.0 mm; Habitus as in Fig. 11 .

Description. Pale yellowish brown on pronotum, elytra and underside surface, head including outer margins of mandibles dark brown except for mouthparts, clypeus and antennae; surface moderately strongly shiny, smooth, glabrous and polish; impunctate, but elytral intervals with tiny, irregularly located punctures, pronotum with puncture-like impressions on and near marginal lateral expanded areas and at central portion of base and front; microsculptural engraved meshes unclear.

Head stout and broad, slightly longer than wide, HL/HW=1.12; eyes rather small but strongly prominent; frons less convex than vertex; clypeus bisetose; labrum 6-setose, frontal margin almost straight; ligula wide, quadrisetose at apex; palps moderately developed; mentum without median tooth, mentum and submentum each with pair of long setae. Antennae short, extended to about basal 1/5 of elytra; pubescent from basal 1/3 of antennomere 4; antennomere 3 as long as 4 and 5 respectively.

Pronotum transverse, PW/HW=1.44–1.47; widest at about middle, PW/PL=1.71–1.72, disc less convex than other species, fore angles broadly round off, and hind angles more or less angular though obtuse; base beaded on median portion, front unbeaded, lateral expanded margins not well-defined, narrow, hardly reflexed; fore and hind transverse impressions shallow but distinct, basal foveae small, median line distinct.

Elytra elongate-ovate, EL/EW=1.44–1.50; moderately convex, base completely bordered, shoulders broadly square; more or less parallel-sided at middle, widest at about middle; striae deep, intervals convex, and subequal in width at middle; apex gently and obliquely truncate, inner angle not denticulate; anterior dorsal pore of interval 3 at about 1/8 of elytra from base, posterior pores at about 1/7 of elytra from apex.

Legs moderate; middle and hind coxae smooth and glabrous in median portion; hind femora moderately expanded, with 2 setae posteriorly; fore tibiae distinctly dilated at apical portions, apical outer angle sharply angular though not protruding, apical margin sinuate, outer margin distinctly serrate; hind tibiae slender, hardly dilated at apex, apical spurs long and sword-like; fore tarsi as stout as middle ones, slightly stouter than hind tarsi; hind tarsomeres 1 distinctly longer than 2, and hind tarsomeres 3 as long as 4 which is deeply and asymmetrically bilobed (with the outer lobe more developed); fore and middle tarsal claws strongly pectinate, hind claws moderately pectinate.

Etymology. This peculiar species is named after Hua Mulan , a beautiful and valorous girl in Chinese legend, referring to the beautiful species in which all of the type series are females.

Distribution. Malaysia ( Fig. 20 i).

Figures

FIGURE 20. Distribution of Hexachaetus species (square: angulatus species group; round: lateralis species group; star: laevissimus species group; heart: vietnamensis species group; triangle: mulan species group; a. H. lateralis lateralis; b. H. lateralis maindroni; c. H. maculatus; d. H. kirschenhoferi n. sp.; e. H. brunki n. sp.; f. H. laevissimus; g. H. vietnamensis n. sp.; h. H. prodigus; i. H. mulan n. sp.)

FIGURES 9 – 11. Habitus of Hexachaetus species (9. H. vietnamensis n. sp.; 10. H. prodigus Tian & Deuve, 2006; 11. H. mulan n. sp. Scale bar: 5.0 mm)