Toliaranella griswoldi, Lorenz & Loria & Harvey & Harms, 2022

Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S. & Harms, Danilo, 2022, The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots, Arthropod Systematics & amp; Phylogeny 80, pp. 649-691 : 649

publication ID

https://dx.doi.org/10.3897/asp.80.e90570

publication LSID

lsid:zoobank.org:pub:AD0B1CC2-6241-4460-B6B8-88E6B95F20D7

persistent identifier

https://treatment.plazi.org/id/8415D05F-4BBF-5412-AE4B-F1FE8C7DC4CB

treatment provided by

Arthropod Systematics & Phylogeny by Pensoft

scientific name

Toliaranella griswoldi
status

sp. nov.

Toliaranella griswoldi sp. nov.

Type material examined.

MADAGASCAR: Toliara Province [Anosy Region]: Holotype ♂ (CAS 9071500 [BLF4810]), allotype ♀ (ZMH-A0016739), paratypes: 11 ♂, 1 ♀ (CAS 9071500 [BLF4810]), 4 ♂, 2 ♀ (ZMH-A0016740 - ZMH-A0016745), Parc National d’Andohahela, Forêt de Manantalinjo, 24°49′1″S 46°36′36″E, 12-16 January 2002, B.L. Fisher, Griswold et al.

Additional material examined.

MADAGASCAR: Toliara Province [ Anosy Region ]: 4 ♂, 4 ♀, 9 nymphs (CAS 9071509 [BLF4984]), Parc National d’Andohahela, Forêt d’Ambohibory, 24°55′48″S 46°38′44″E, 16-20 January 2002, Fisher, Griswold et al GoogleMaps .; 1 ♂, 1 ♀ (CAS 9071516 [BLF4815]), Parc National d’Andohahela, Foret de Manantalinjo , 24°49′1″S 46°36′36″E, 12-16 January 2002, Fisher, Griswold et al GoogleMaps .

Diagnosis.

Differs from T. pumila sp. nov. in different cheotaxy: est on fixed finger situated between sb and t of the movable finger on sagittal level (est on same sagittal level with sb in T. pumila sp. nov.); most compact carapace of the southern feaellid members (1.27/1.23 ♂/♀ times longer than broad versus 1.31-1.37/1.31-1.39 ♂/♀).

Etymology.

This species is a patronym honoring Charles Griswold who collected many Malagasy feaellids.

Description.

The following description is based on holotype and allotype. - Carapace (Fig. 22A View Figure 22 ): Very compact, heavily granulate, 1.20-1.31 (♂), 1.18-1.31 (♀) times longer than broad. - Pedipalps (Fig. 22C, D View Figure 22 ): Similar trichobothrial pattern as T. pumila sp. nov. but the following difference: est on fixed finger situated between sb and t of the movable finger on sagittal level. Chelal fixed finger with 9-10 chelal teeth in the OR, 12-14 in the MR and 9-11 in the IR; movable finger with 6-8 in the OR, 7-11 in the MR and 7-8 in the IR. Fixed finger with 6-7 terminal teeth equal in size and one large one, movable finger with 7-8 equally sized terminal teeth. - Dimensions (mm): Holotype ♂: Body length 1.62; abdomen length 1.13, width (with pleura) 1.05, width (without pleura) 0.91; carapace length 0.46, width 0.35. Pedipalp: trochanter 0.22; femur length 0.49, width 0.27; patella 0.35; chela (without pedicel) 0.35; hand length 0.10, width 0.11; movable finger 0.26. Leg I: trochanter 0.07; femur 0.14; patella 0.13; tibia 0.11; tarsus 0.21. Leg IV: trochanter 0.13; femur 0.10; patella 0.21; tibia 0.20; tarsus 0.26. Allotype ♀: Body length 2.04; abdomen length 1.47, width (with pleura) 1.33, width (without pleura) 1.11; carapace length 0.53, width 0.45. Pedipalp: trochanter 0.25; femur length 0.59, width 0.35; patella 0.46; chela (without pedicel) 0.44; hand length 0.10, width 1.14; movable finger 0.35. Leg I: trochanter 0.11; femur 0.16; patella 0.14; tibia 0.16; tarsus 0.25. Leg IV: trochanter 0.18; femur 0.14; patella 0.27; tibia 0.31; tarsus 0.30.

Variations.

Body length 1.54-1.67 (♂), 1.96-2.24 (♀); abdomen length 1.08-1.18 (♂), 1.41-1.63 (♀); width (with pleura) 0.98-1.09 (♂), 1.27-1.41 (♀); width (without pleura) 0.84-0.95 (♂), 1.10-1.19 (♀); 1.20-1.29 (♂), 1.27-1.37 (♀) times longer than broad; carapace length 0.42-0.48 (♂), 0.53-0.57 (♀); width 0.35-0.38 (♂), 0.41-0.45 (♀); 1.20-1.31 (♂), 1.18-1.29 (♀) times longer than broad. Pedipalp: trochanter 0.19-0.22 (♂), 0.24-0.26 (♀); femur length 0.40-0.49 (♂), 0.54-0.59 (♀); width 0.25-0.28 (♂), 0.33-0.37 (♀); 1.60-1.81 (♂), 1.53-1.69 (♀) times longer than broad; patella 0.33-0.36 (♂), 0.45-0.48 (♀); chela (without pedicel) 0.35-0.38 (♂), 0.43-0.47 (♀); hand length 0.08-0.11 (♂), 0.10-0.11 (♀); width 0.11-0.13 (♂), 0.13-0.14 (♀); 0.73-0.91 (♂), 0.71-0.79 (♀) times longer than broad; movable finger 0.26-0.30 (♂), 0.35-0.37 (♀).

GenBank Code.

OP589959, OP589960. The species differs from all congeners by more than 3.7% in the H3 dataset.

Habitat.

Specimens were found in sifted litter of spiny forest thicket and tropical dry forest at an elevation of 150-300 m.

Distribution.

Currently known from the type locality (BLF4810) and two additional localities of (BLF4984 and BLF4815) in the Anosy Region (former Toliara Province) in southern Madagascar.