Nannaria fritzae, Means & Hennen & Marek, 2021

Means, Jackson C., Hennen, Derek A. & Marek, Paul E., 2021, A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae), ZooKeys 1030, pp. 1-180: 1

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scientific name

Nannaria fritzae

sp. nov.

Nannaria fritzae   sp. nov. Figs 55 View Figure 55 , 56 Vernacular name: "Kathlyn Fritz’s Twisted-Claw Millipede" View Figure 56

Material examined.

Holotype: United States - Georgia • ♂ Chattooga County, hillside by Lake Marvin Rd.; 34.5628°N, - 85.0681°W; elev. 385 m; 5 Mar. 2017; hand collected; J. Means, K. Means leg.; VTEC MPE02359 View Materials . GoogleMaps  

Paratypes: United States - Georgia • 1 ♂; same collection data as holotype; VTEC MPE02377 View Materials GoogleMaps   1 ♂; same collection data as holotype; VMNH MPE02378 View Materials GoogleMaps   .


Adult males of Nannaria fritzae   sp. nov. are distinct from other Nannaria   and the nearby Nannaria   sp. nov. ‘Amicolola’ ( Nannaria wilsoni   species group) based on the following combination of characters. Gonopods. Gonopodal acropodite gently curving medially throughout, not straight with abrupt medial bend at apex as in Nannaria   sp. nov. ‘Amicolola.’ Distal zone short, rounded, not serpentine as in Nannaria   sp. nov. ‘Amicolola.’ Acropodite with small, tooth-like medial flange near apex (Fig. 55A View Figure 55 , red arrow). Prefemur with long acicular prefemoral process bent at base and directed medially-not curving laterally, crossing acropodite, as in N. minor   or not large, laminate as in Nannaria   sp. nov. ‘Amicolola.’ Prefemoral spine reduced to thorn-like structure, arising from prefemoral process, directed ventrally (Fig. 55B View Figure 55 , red triangle)-not curving cephalically as in N. minor   . Telopodite basal zone ca. 1/2 length of acropodite, not < 1/2 as in N. minor   or > 1/4 as in Nannaria   sp. nov. ‘Amicolola.’ Color. Tergites with orange paranotal spots (Fig. 56 View Figure 56 ), reduced pigmentation of paranotal spots of terminal segments. Light brown background. Dorsum of collum smooth with orange and white margin.


♂ holotype (VTEC, MPE02359): BL = 28.2, CW = 3.8, IW = 2.3, ISW = 0.8, B11W = 4.6, B11H = 3.0.


No known variation.


Known only from the type locality (Georgia: Chattooga County; Suppl. material 7; Fig. 126 View Figure 126 ). Distribution area: N/A; status: MRE.


Individuals of N. fritzae   sp. nov. were collected on a hillside in a mesic hardwood forest dominated by oak and maple. Specimens were found under leaf litter in a dry creek bed near the base of an oak tree.


This species is named for its co-collector, Kathlyn Fritz Means. The name is a genitive noun derived as a matronym.

Type locality.

United States, Georgia, Chattooga County, hillside by Lake Marvin Rd.; 34.5628°N, - 85.0681°W.