Tetragonoschema (Tetragonoschema) barriesi, Bílý, Svatopluk, 2012

Bílý, Svatopluk, 2012, A revision of the genera Anilaroides Théry, 1934, stat. nov. and Tetragonoschema Thomson, 1857 (Coleoptera: Buprestidae: Buprestinae: Anthaxiini), Zootaxa 3521, pp. 1-38 : 16-17

Tetragonoschema (Tetragonoschema) barriesi sp. nov.

( Figs. 44 , 61 , 76 )

Specimens examined. Holotype (male): “ Paraguay, Prov. Boqueron, 9.–11.xii.2010, NP Enciso , 257 m, 21°12ʹS 61°39ʹW, Sv. Bílý leg.” ; allotype (female): the same data ; paratypes: the same data (11 males, 22 females) ; “ Paraguay, Prov. Boqueron, 5.–7.xii.2010, Loma Plata, 141 m, 22°23ʹS 59°50ʹW, Sv. Bílý leg.” (4 females) ; “ Paraguay, Chaco alto, Prov. Boquerón, Enciso NP, 257 m, 21°12ʹ38ʹ S 61°39ʹ26ʹ W, 9.–12.xii.2010, Barries, Bílý & Cate leg.” (8 males, 10 females) ; “ Paraguay, Chaco médio, Prov. Boquerón, Loma Plata env., 141 m, 22°23ʹ18ʹ S 59°50ʹ0 6ʹ W, 5.–15.xii.2010, Barries, Bílý & Cate leg.” (7 males, 4 females) . Holotype and allotype deposited in NMPC, paratypes in NMPC and WBCW.

Diagnosis. Medium-sized (3.6–5.2 mm), subparallel, rather flattened, matt; entire dorsal surface dark bronze with a red lustre, ventral surface bronze in male, dark bronze with blue reflections in female; dorsal surface asetose, frons with extremely short, microscopic, white pubescence; ventral surface with very short, sparse, white pubescence.

Description of the male holotype. Head relatively small, somewhat retracted into pronotum, narrower than anterior pronotal margin; frons widely, deeply grooved, supra-antennal prominences well-developed; vertex slightly convex, about twice as wide as width of eye; eyes small, reniform, not projecting beyond outline of head; sculpture of head consisting of very dense, small, polygonal cells with flat central granules; antennae short, barely reaching midlength of lateral pronotal margins when laid alongside the pronotum; scape claviform, slightly curved, 3 times as long as wide, pedicel oval, 1.5 times as long as wide; third antennomere subcylindrical, 1.6 times as long as wide, antennomeres 4–5 obtusely triangular, 1.4 and 1.2 times as long as wide, antennomeres 6–10 trapezoidal, about as long as wide; terminal antennomere ovoid, slightly longer than wide.

Pronotum moderately convex, 1.70 times as wide as long, with two small, rounded, discal impressions; anterior margin deeply biarcuate, medial lobe rather produced, posterior margin biarcuate; lateral margins distinctly angulate, rectilinearly converging to anterior angles from widest point, with obtuse-angled posterior angles, maximum pronotal width at anterior two fifths; lateroposterior depressions wide but shallow; pronotal sculpture homogenous, consisting of small, dense, polygonal cells with small central granules only along lateral margins; lateral carina short, barely reaching midlength of lateral margins, not visible from above; posterior angles with prehumeral keel (as in the genus Agrilus Curtis, 1825 ) reaching midlength of lateral margins ( Fig. 61 ). Scutellum small, triangular, nearly twice as long as wide, microsculptured.

Elytra flattened, 1.16 times as long as wide, narrowed behind the humeri, strongly uneven with several deep depressions: one wide, common, transverse depression at midlength, each elytron with a wide, triangular, preapical depression and with a deep, wide, elongate, lateral depression at posterior two thirds; transverse, humeral depression deep and wide, completely interrupted by a large, blister-like elevation near the humeral callosity; humeral callosities well-developed; elytral epipleura wide, reaching elytral suture, easily visible from above; elytral sculpture dense, rugose with very indistinct punctation on basal half, simply rugose with small, dense granules on posterior half; central portion of elytral disc more lustrous, with weakly developed, fine, oblique punctation; apex of each elytron obtusely truncate, both outer and sutural angles widely rounded.

Ventral surface shiny, finely ocellate; prosternal process flat, wider than long, slightly enlarged behind the procoxae, sharply pointed apically; anal ventrite angulately produced apically, with wide, deep depression along lateral sides. Legs relatively long, slender, protibiae slightly curved, meso- and metatibiae straight, simple; all tarsi much shorter than tibiae; tarsal claws short, slender, hook-shaped, only weakly enlarged at base.

Aedeagus ( Fig. 76 ) elongate, parameres widened at basal third, with small, sharp, preapical spines; median lobe sharply pointed apically.

Sexual dimorphism. Female differs from male in the somewhat stouter body, dark bronze frons (frons with a red lustre in male) and in the blue lustre of the ventral surface (bronze in male).

Variability. Except for the size (3.6–5.2. mm) no substantial variation was observed.

Measurements. Length: 3.6–5.2 mm (holotype 4.0 mm); width: 1.6–2.4 mm (holotype 1.9 mm).

Bionomy. Unknown.

Etymology. Tetragonoschema (T.) barriesi sp. nov. is named after my friend and colleague Wolfgang Barries (Vienna), a specialist on the genus Chrysobothris Eschscholtz, 1829 .

Distribution. Paraguay (prov. Boquerón).

Differential diagnosis. Tetragonoschema (T.) barriesi sp. nov. belongs to the group of species with welldeveloped, wide elytral epipleura which are easily visible from above and with parameres with a preapical spine ( T. (T.) albopilosum and T. (T.) rubromarginatum ). Except for the characters given in the key it differs from T. (T.) rubromarginatum in its colouration and different shape of anal ventrite and male genitalia ( Figs. 76 vs. 75); from T. (T.) strandi it differs in its less convex body, very short frontal pubescence, different male genitalia ( Figs. 76 vs. 77) and also in the dark colouration.

Figures

FIGURES 37 – 45. 37 – 40. Historical types: 37 — Tetragonoschema fallaciosum Théry, 1944, paratype, 5.5 mm; 38 — T. santafeanum Obenberger, 1947, lectotype, 4.2 mm; 39 — T. vianai Obenberger, 1947, lectotype, 4.1 mm; 40 — T. monrosi Cobos, 1949, holotype, 4.5 mm. 41 – 45. Habitus: 41 — Anilaroides brasiliensis (Kerremans, 1897), male, 4.3 mm (Brasil, S. Antonio da Barra); 42 — T. (Patagoschema) cobosi sp. nov., male, 2.7 mm (paratype); 43 — T. (Tetragonoschema) aeneum Kerremans, 1899, male, 4.5 mm (Paraguay, Serro Acahay); 44 — T. (T.) barriesi sp. nov., male, 5.0 mm (paratype); 45 — T. (T.) cupreocingulatum Hoscheck, 1931, male, 3.7 mm (Paraguay, Loma Plata).

FIGURES 55 – 63. 55 – 57. Habitus: 55 — Tetragonoschema (T.) trinidadense Bellamy 1991, male, 4.5 mm, typical form (Trinidad, S. Benedico); 56 — the same, male, 3.5 mm, blue form (Venezuela, Guamacho); 57 — the same, female, 5.0 mm, green form (Venezuela, Maicillal); 58 – 63. 58 — elytral apex of Tetragonoschema (Patagoschema) torresi Cobos, 1959 (holotype); 59 — the same, T. (T.) humerale Waterhouse, 1882 (Mexico, Michoacán); 60 — the same, T. (T.) strandi Obenberger, 1924 (lectotype); 61 — pronotum of T. (T.) barriesi sp. nov. (holotype), lateral view; 62 — the same, T. (T.) strandi (holotype); 63 — anal ventrite of T. (P.) cobosi sp. nov. (holotype).

FIGURES 72 – 81. Aedeagi: 72 — Tetragonoschema (Patagoschema) patagonicum Obenberger, 1922 (holotype); 73 — T. (P.) torresi Cobos, 1959 (holotype); 74 — T. (P.) cobosi sp. nov. (holotype); 75 — T. (T.) rubromarginatum Théry, 1944 (lectotype); 76 — T. (T.) barriesi sp. nov. (holotype); 77 — T. (T.) strandi Obenberger, 1924 (Argentina, Tucuman); 78 — T. (T.) cupreocingulatum Hoscheck, 1931 (lectotype); 79 — T. (T.) fossicolle Kerremans, 1900 (lectotype); 80 — T. (T.) humerale Waterhouse, 1882 (Mexico, Michoacan); 81 — T. (T.) trinidadense Bellamy, 1991 (holotype).