Anisotoma, Panzer, 1797

Peck, Stewart B. & Newton, Alfred F., 2017, An Annotated Catalog of the Leiodidae (Coleoptera) of the Nearctic Region (Continental North America North of Mexico), The Coleopterists Bulletin 71 (2), pp. 211-258 : 235-237

Genus ANISOTOMA Panzer, 1797

Anisotoma Panzer, 1797: 8 . Type species: Tritoma glabra Fabricius, 1787 (designated by Hatch, 1929b: 50; “ A. picea (Illig) ” designated by Thomson 1859: 58).

Note. Authority given as Knoch in Kugelann (1794) (Neave 1939–49); two original species (“ picea Kug. ”, “ glabra Kug. ”).

North American Revision. Wheeler (1979) and Wheeler and Miller (2005).

Biology. The species generally eat plasmodia or fruiting bodies of Myxomycetes slime molds (e.g., Wheeler 1979; Lawrence and Newton 1980; Newton 1984; Wheeler 1984a, b).

Distribution. Holarctic, Nepal, China, Taiwan, N Neotropical (to Panama) (57 species). New (unpublished) state and province records are included in the distributions below based on SBP voucher specimens deposited in CMNC.

=Pentatoma Schneider, 1792: 339 (preoccupied, not Olivier, 1789). Type species: Sphaeridium humerale Olivier, 1790 (cited as Fabricius, 1792) (designated by Hatch 1929b: 50).

= Anisotoma Illiger, 1798: 69 (attributed to Knoch). Type species: none designated? (13 original species, including “ piceum Panzer ” and “ glabrum Panzer”).

= Leiodes ; Schmidt 1841: 132 (misidentification of Leiodes Latreille, 1797 ).

= Eucyrta Portevin, 1927: 82 (preoccupied, not Felder 1874); Wheeler 1979: 267 (synonymy). Type species: Eucyrta didymata Portevin, 1927 (designated by Hatch, 1929b: 57).

A. amica Brown, 1937b: 201 ; Hatch 1957: 31; Wheeler 1979: 293. Holotype male in CNCI. Type locality: Mt. Layall, Quebec. North American distribution: Canada: AB, BC, MB, NF, ON, QC. USA: AK, AZ, CO, NM, WA. Biology: Fungal associates ( Stephenson et al. 1994).

A. basalis ( LeConte, 1853: 285) (Leiodes) ; Horn 1880: 299 ( Liodes ); Brown 1937b: 200; Wheeler 1979: 292; Majka and Langor 2008: 382. Lectotype female in MCZC (designated by Wheeler 1979: 292). Type locality: Pennsylvania or New Jersey. North American distribution: Canada: MB, NB, NF, NS, ON, QC. USA: AL, AR, CT, GA, IL, IN, KY, LA, MA, MD, ME, MI, MN, MS, NC, NH, NJ, NY, OK, PA, RI, SC, TX, VA, VT, WI. Larva: Wheeler (1990a, b). Biology: Fungal associates (Wheeler and Hoebeke 1990; Stephenson et al. 1994).

= Leiodes dichroa LeConte, 1853: 285 : Wheeler 1979: 292 (synonymy). Lectotype male in MCZC (designated by Wheeler 1979: 292). Type locality: Ann Arbor, Michigan.

A. bifoveata Wheeler, 1979: 273 . Holotype male in MCZC. Type locality: Conkles Hollow , Hocking County, Ohio. North American distribution: Canada: ON. USA: AL, AR, CT, GA, IL, IN, KY, LA, MI, MO, MS, NC, NY, PA, SC, TX, VA, WV.

A. blanchardi ( Horn, 1880: 295) (Liodes) ; Brown 1937b: 196; Wheeler 1979: 273; Majka and Langor 2008: 383. Lectotype male in MCZC (designated by Wheeler 1979: 273). Type locality: Tewksbury, Massachusetts. North American distribution: Canada: MB, NB, NS, ON, QC. USA: AL, FL, GA, IL, KY, LA, MA, MD, ME, MI, MN, NC, NH, OH, PA, SC, TN, TX, VA, VT. Larva: Kilian (2005a). Biology: Fungal associates ( Stephenson et al. 1994).

A. confusa ( Horn, 1880: 299) (Liodes) ; Brown 1937b: 197; Hatch 1957: 31; Wheeler 1979: 288. Holotype male in MCZC. Type locality: “Western Nevada” (probably vicinity of Lake Tahoe, Reno, or Carson City). North American distribution: Canada: AB, BC. USA: AK, CA, OR, NV, WA.

= A. interstrialis Hatch, 1936: 38 ; Brown 1937b: 197; Wheeler 1979: 288 (synonymy). Holotype in USNM. Type locality: Seattle, Washington.

A. discolor (F. E. Melsheimer, 1844: 103) (Leiodes) ; LeConte 1853: 285; Horn 1880: 298; Brown 1937b: 202; Wheeler 1979: 295; Majka and Langor 2008: 384. Lectotype female in MCZC (designated by Wheeler 1979: 295). Type locality: “Pennsylvania.” North American distribution: Canada: BC, MB, NB, NS, ON, QC. USA: AL, AR, CT, DC, FL, GA, IA, IL, IN, KY, LA, MA, MD, ME, MI, MN, MO, MS, NC, NH, NJ, NY, OH, OK, PA, RI, SC, TN, TX, VA, WV. Larva: Kilian (2003).

= Anistoma [sic] piceum F. V. Melsheimer, 1806: 12 (attributed to Knoch; nomen nudum); F. E. Melsheimer 1844: 103 (as synonym of Leiodes discolor F. E. Melsheimer ); Wheeler 1979: 288 (synonymy).

A. errans Brown, 1937b: 202 ; Hatch 1957: 32; Wheeler 1979: 277; Majka and Langor 2008: 384. Holotype male in CNCI. Type locality: Biscotasing, Ontario. North American distribution: Canada: AB, BC, NB, NF, NS, ON, QC, SK, YT. USA: AK, CA, ID, ME, MI, NH, OR, RI, VT, WA, WY. Biology: Fungal associates ( Stephenson et al. 1994). Larva: Newton (1991).

A. expolita Brown, 1937b: 199 (replacement name for Leiodes polita LeConte, 1853 ); Wheeler 1979: 300. North American distribution: USA: AR, IL, IN, KY, PA, SC.

= Leiodes polita LeConte, 1853: 285 (preoccupied, not Silpha polita Marsham, 1802 , not now congeneric but replaced before 1961 and replacement name in use ( ICZN 1999: Art. 59.3)); Horn 1880: 297 ( Liodes ). Holotype male in MCZC. Type locality: “ South Carolina.”

A. geminata ( Horn, 1880: 299) (Liodes) ; Brown 1937b: 196; Wheeler 1979: 288; Majka and Langor 2008: 385. Lectotype male in MCZC (designated by Wheeler 1979: 288). Type locality: Tyngsboro, Massachussets. North American distribution: Canada: BC, NB, NF, NS, ON, QC. USA: CT, GA, IL, IN, KY, MA, ME, MI, NC, NH, NJ, NY, OH, PA, RI, SC, TN, TX, VA, WI. Biology: Fungal associates ( Stephenson et al. 1994).

A. globososa Hatch, 1929b: 56 (replacement name for A. globosa ( LeConte, 1850)) ; Hatch 1957: 31; Brown 1937b: 197; Wheeler 1979 a: 298; Majka and Langor 2008: 386. North American distribution: Canada: AB, BC, MB, NF, ON, QC, SK. USA: AK, CO, CT, ID, MA, ME, MI, NH, SD, TX, WA, WI. Biology: Fungal associates ( Stephenson et al. 1994).

= Cyrtusa globosa LeConte, 1850: 222 (preoccupied in Anisotoma , not Tetratoma globosa Herbst, 1791 ; not now congeneric, but replaced before 1961 and replacement name in use ( ICZN 1999: Art. 59.3)); LeConte 1853: 285 ( Leiodes ); Horn 1880: 297 ( Liodes ). Lectotype male in MCZC (designated by Wheeler 1979: 298). Type locality: Sault Ste Marie, Michigan.

A. humeralis ( Olivier, 1790: 15) (Sphaeridium) ; Hatch 1929b: 51; Wheeler 1979: 290. Type in MNHN? Type locality: “environs de Paris”. North American distribution: Canada: BC. USA: IN. Extra-limital distribution: Europe; a Holarctic species. Larva: Ratajczak (1995) and Kilian (1998).

A. inops Brown, 1937b: 198 ; Wheeler 1979: 299; Majka and Langor 2008: 386; Bouchard 2001: 130. Holotype male in CNCI. Type locality: Arnprior, Ontario. North American distribution: Canada: NB, NF, NS, ON, QC. USA: ME, NC, NH, NY.

A. nevadensis Brown, 1937b: 199 (replacement name for Liodes globosa var. bicolor Horn, 1880 ); Hatch 1957: 31; Wheeler 1979: 300. North American distribution: USA: CA, NV, OR, WA.

= Liodes globosa var. bicolor Horn, 1880: 297 (preoccupied by Anisotoma bicolor Schmidt, 1841 ). Lectotype male in MCZC (designated by Wheeler 1979: 300). Type locality: “ Nevada ” [probably vicinity of Lake Tahoe, Reno, or Carson City].

A. obsoleta ( Horn, 1880: 298) (Liodes) ; Brown 1937b: 201; Majka and Langor 2008: 386 (revalidation of name). Lectotype male in MCZC (designated by Wheeler 1979: 276). Type locality: none given, range stated as “New Hampshire to Virginia and Canada.” North American distribution: Canada: BC, MB, NB, NF, NS, ON, QC. USA: AL, AR, CT, FL, GA, IA, IL, KY, LA, MA, ME, MI, MN, MS, NC, NH, NY, OH, PA, SC, TN, VA, VT, WI, WV. Biology: Fungal associates Stephenson et al. 1994). Larva: Kilian (2005b) (as A. horni Wheeler ).

= A. horni Wheeler, 1979: 276 (unnecessary replacement name for Liodes obsoleta Horn, 1880 , incorrectly thought to be preoccupied by Pallodes obsoletus F. E. Melsheimer, 1844 (now in Anogdus LeConte )). Biology: Bouchard (2001).

A. reticulonota Wheeler, 1979: 298 . Holotype male in MCZC. Type locality: White Mountains , Apache County, Arizona. North American distribution: USA: AZ, NM.