Anaclastoctedon ancistrodes, Plant, 2010

Plant, Adrian R., 2010, Anaclastoctedon (Diptera: Empididae: Hemerodromiinae), A New Genus From Asia And Australia, Raffles Bulletin of Zoology 58 (1), pp. 15-25 : 20-21

Anaclastoctedon ancistrodes , new species

( Fig. 9 )

Material examined. – Holotype. Male , AUSTRALIA: NSW, Blue Mountains N.P., Blackheath, Govetts Leap [approx. 33°37'S 150°17'E], ex. dry scler. / creek, 4 Apr.1994, coll. B. J. Sinclair ( CNC).

Paratypes. – Same data as holotype: 8 males, 2 females ( CNC) .

Diagnosis. – Black species, larger than A. lek new species and with head only moderately dorsoventrally flattened; distinguished from A. prionton new species primarily by having wings distinctly brownish, a dark halter and distinctive male genitalia.

Description. – Male. Length 2.5mm. Head moderately dorsoventrally compressed, black, face dusted paler. All setae black, lower postocular setae strong with weaker row in front close to eye margin, only minute setulae behind mouth. Proboscis blackish, palpus yellow with black seta apically. Basal antennal segments yellow; postpedicel blackish, yellowish below; stylus 4–5× long as postpedicel.

Thorax black, dusted bluish grey; all setae black, dorsocentrals, postpronotal, upper notopleural and supraalar similarly strong.

Legs brownish yellow, setae black; coxae, F 1 dorsally and tibiae obscurely darker; distal tarsomeres brown. F 1 very stout, about 5 av and 4–5 pv strong black setae between which are rows of about 17 av and 20 pv denticles; dorsal fringe of setae distinct, longest near base. T 2 with strong apicoventral seta.

Abdomen brown, rather paler below; sparsely setate, tergite 5 with fan of strong dark setae. Epandrium and hypandrium ( Fig. 9 ) dark brown, bearing distinct short setae. Epandrium almost circular with short ad lobe bearing regular series of small setulae dorsally. Cercus with major lobe elongate, slightly broader apically, stout spines and strong setae distally; basal lobe smaller, broadened subapically. Parameral sheath broad with two recurved pointed processes subapically and inverted U-shaped apical process. Phallus dark basally, abruptly yellowish distally.

Wing membrane vaguely brownish, veins brown; vein M 2 completely absent. Halter greyish-brown.

Female. – Length 2.6–2.8mm. Similar to male but postpedicel uniformly dark and tergite 5 without fan of strong setae. Cercus brownish.

Etymology. – The specific epithet ancistrodes (Greek) means barbed and refers to the recurved pointed processes on the parameral sheath.

Remarks – This species is described from the Blue Mountains National Park, New South Wales in eastern Australia during April. The habitat at the type locality is predominantly dry sclerophyll forest at about 1,000 m with localised seepages and moisture associated vegetation.

Figures

Figs. 9–12. Anaclastoctedon new genus: 9. A. ancistrodes new species, male terminalia; 10. A. sano new species, male terminalia; 11–12. A. prionton new species; 11. male terminalia; 12. apex of parameral sheath, dorsal aspect.