Bisnius fimetarius (Gravenhorst)

Pietrykowska-Tudruj, Ewa & Staniec, Bernard, 2011, The morphology of the pupae of six species of Philonthina (Coleoptera, Staphylinidae, Staphylinini) with taxonomic remarks, Zootaxa 2865, pp. 53-67 : 56

Bisnius fimetarius (Gravenhorst)

( Figs 1–3 , 41–42, 52, 61, 62)

Body length: 3.57–4.40 mm (mean 4.16 mm); body width: 1.43–1.84 mm (mean 1.67 mm); head width: 0.80–1.24 mm (mean 1.03 mm); pronotum width: 1.11–1.17 mm (mean 1.15 mm), about 1.1 x as broad as long; body moderately stocky and weakly sclerotized. Antennae reaching to about half of elytra length ( Fig. 2 ). Anterior margin of pronotum with 10–14 projections (6/6, 7/7, 7/7, 5/5, 7/5, 6/5, 6/5 projections on sides in individuals). Wings almost reaching the posterior margin of 1st (morphologically 3rd) clearly visible abdominal segment ( Figs 1, 2 ). Each fore, middle, and hind tibiae with 5, 8–10, and 6 protuberances respectively. Tarsi of hind legs slightly protruding from anterior margin of 4th (morphologically 6th) clearly visible abdominal segment ( Figs 1, 2 ). Abdomen moderately and gradually widened from segment II to IV and narrowed to the terminal part of the body ( Fig. 3 ). Abdominal tergites with microstructure as in G. osseticus ( Fig. 19 A). Abdominal segments VII and VIII each bearing a pair of setiform projections on sides, projections similar in length and structure to these one in G. osseticus ( Figs 19, 20 ). Terminal sternite of female pupa with two well-developed ventral prolongations (Fig. 41); in male pupa sternite IX as in Fig. 42; ventral and terminal prolongation with numerous, sharp, cuticular processes (Figs 41A, B). The first pair of functional spiracles elongated, gradually narrowed apically, almost as long as wide ( Fig. 52 ); atrophied spiracles as in Figs 61, 62 .

Figures

FIGURES 1 – 6. Pupa of Bisnius fimentarius (1 – 3) and Gabrius osseticus (4 – 6). (1, 4) Ventral aspect, (2, 5) lateral aspect, (3, 6) dorsal aspect.

FIGURES 19 – 40. Pupa of Gabrius osseticus (19, 20), Philonthus atratus (21 – 25), P. carbonarius (26 – 32), P. debilis (33 – 35) and P. decorus (36 – 40). (19, 21, 26, 27, 36) Lateral part of segment VII with microstructure (19 A, 26 A) and spine (27); (20, 22, 33, 39, 40) lateral part of segment VIII with microstructure (39 A) and spine (40); (23, 24, 28, 29) lateral part of segment IV with spine (24, 29); (25, 31, 32, 34, 35, 37, 38) lateral part of segment V with microstructure (34 A) and spine (32, 35, 38).

FIGURES 52 – 70. Pupa of Bisnius fimentarius (52, 61, 62), Gabrius osseticus (53, 58, 63, 64), Philonthus atratus (54, 59, 65), P. carbonarius (55, 66), P. debilis (56, 60, 68 – 70) and P. d e c o r u s (57, 67). (52 – 60) The first pair of functional spiracles; (61 – 70) atrophied spiracles.