Sabaconidae Dresco, 1970

SCHÖNHOFER, AXEL L., 2013, Axel L. Schönhofer, Zootaxa 3679 (1), pp. 1-68 : 16-19

publication ID

https://doi.org/ 10.11646/zootaxa.3679.1.1

publication LSID

lsid:zoobank.org:pub:770B1AA2-869F-4F1B-8010-0E3FAECB0592

persistent identifier

https://treatment.plazi.org/id/7E65412B-FE57-FFC8-FF2C-FC6FE344FA96

treatment provided by

Felipe

scientific name

Sabaconidae Dresco, 1970
status

 

Family Sabaconidae Dresco, 1970 View in CoL

Sabaconidae Dresco, 1970: 1210 View in CoL (only including Sabacon View in CoL ); type genus Sabacon Simon, 1879a View in CoL by monotypy and original designation;— Martens 1978: 225; 1983: 269; Shear 1986: 3 (compr. Sabacon View in CoL , Taracus ); Giribet et al. 2010: 421 (compr. Hesperonemastoma View in CoL , Sabacon View in CoL , Taracus ); new composition (compr. Sabacon View in CoL ).

Diagnosis. Dyspnoi of the superfamily Ischyropsalidoidea with a disjunct Holarctic distribution. Defined by peculiarities of male genital morphology, molecular phylogenetic evidence and the peculiar shape and functionality of palps. Palps densely set with bottlebrush-shaped hairs, inflated in all parts, most pronounced in females and juveniles; tarsus freely movable to dorsal and ventral side. Chelicerae small to moderate in size, in many species males with glandular fields on dorsal side of proximal cheliceral segment, often associated with large apophyses. Truncus penis of complex shape, with a deeply split base drawn out in two long roots, forming an upside-down Ystructure with the two basal branches bent towards the truncus. A single muscle situated outside the truncus and mounted between the truncus roots, connected with the distal part of the penis by a long tendon running through the truncus. Further characters of truncus, glans and stylus very variable, but penial stylus generally very long, if not modified. Ventral region poorly sclerotised, generally smooth and glossy without armature or microstructure. Labium absent, coxae without coxal apophyses. Frontal margin of opisthosoma with a median furrow. Two metapeltidial sensory cones.

Genus Sabacon Simon, 1879a (35 species, 4 subspecies)

Sabacon Simon, 1879a: 266 View in CoL ; type species Sabacon paradoxus Simon, 1879a View in CoL by monotypy.

Phlegmacera Packard, 1884: 203 View in CoL ; type species Phlegmacera cavicolens Packard, 1884 View in CoL , by monotypy;—syn. Roewer 1914: 123.

Tomicomerus Pavesi, 1899: 533 View in CoL ; type species Tomicomerus bryantii Banks, 1898 View in CoL by monotypy;—syn. Shear 1986: 5 (see also Remarks on bryantii View in CoL ).

Parasabacon Hansen & Sørensen, 1904: 83 ; type species Nemastoma crassipalpis L. Koch, 1879 by monotypy;—syn. Roewer 1914: 123.

Remarks. Roewer (1914) and subsequent authors treated Sabacon as neuter in gender, while Gruber (2003) clarified it to be masculine, but did not list species names with corrected ending. In a list of species from France and Spain, Prieto (2008) stated that Gruber had corrected the endings. Additional mandatory changes made for the first time according to Gruber are indicated by “new ending”.

Etymology. Gender masculine; from Greek “Sabákon” (Egyptian king). For details see ( Gruber 2003).

akiyoshiensis Suzuki, 1963 ; loc. typ.: Japan, Yamaguchi Pref., Miné-shi, Ōmine-machi, Satoyamase, Satoyamaseno-ana Cave.

Sabacon akiyoshiensis Suzuki, 1963: 156 View in CoL

Sabacon akiyoshiense — Suzuki 1974a: 102.

altomontanus Martens, 1983 View in CoL ; loc. typ.: France, Dép. Hautes-Pyrénées, SW Lannemézan.

Sabacon altomontanum Martens, 1983: 290 View in CoL .

Sabacon altomontanus View in CoL — Prieto 2008: 51.

astoriensis Shear, 1975: 24; loc. typ.: USA, Oregon, Clatsop Co., Fort Stevens State Park.

briggsi Shear, 1975: 26; loc. typ.: USA, California, Marin Co., Bolinas Ridge.

bryantii ( Banks, 1898) View in CoL ; loc. typ.: USA, Alaska, Mt. St. Elias, Malaspina Glacier. Remarks. Changes to the original spelling “ bryantii View in CoL ” are prohibited (ICZN 33.4). The generic placement of bryantii View in CoL remains to be investigated. It appears to have morphological traits of both Sabacon View in CoL and Taracus , but new material is needed for further investigation ( Shear 1975, 1986).

Phlegmacera bryantii Banks, 1898: 16 View in CoL .

Tomicomerus bryantii View in CoL — Roewer 1914: 127.

Tomicomerus bispinosus Pavesi, 1899: 533 View in CoL ;—syn. Roewer 1914: 127.

Tomicomerus bryanti — Shear 1975: 26 (incorrect subsequent spelling).

Sabacon bryanti — Shear 1986: 5.

cavicolens ( Packard, 1884) View in CoL ; loc. typ.: USA, Kentucky, Carter Co., Bat Cave.

Phlegmacera cavicolens Packard, 1884: 203 View in CoL .

Phlegmacera cavicoleus — Banks 1894b: 52; Banks 1901: 677 (misspellings).

Nemastoma inops Packard, 1884: 203 ;—syn. Shear 2010b: 107.

Sabacon spinosus Weed, 1893: 575 View in CoL ;—syn. Banks 1894b: 40.

Sabacon jonesi Goodnight & Goodnight, 1942: 1 View in CoL ; syn. n. Remarks. Shear (1975: 8) tentatively placed jonesi View in CoL close to cavicolens View in CoL which is accepted and the synonymy formally established here.

Hesperonemastoma inops View in CoL — Gruber 1970: 138.

Sabacon cavicolens View in CoL — Shear 1975: 13.

chomolongmae Martens, 1972: 312; loc. typ.: Nepal, Khumbu, Dhud-Kosi Valley, Lughla.

crassipalpis (L. Koch, 1879) ; loc. typ.: Siberia, Tunguska, 4 mi S om Podsk (see also Martens 1989: 370). Nemastoma crassipalpis L. Koch, 1879: 111 .

Sabacon crassipalpis View in CoL — Roewer 1914: 125; Martens 1989: 370 (redescr.); Chemeris & Logunov 2000: 207 (redescr.).

dentipalpis Suzuki, 1949 View in CoL ; loc. typ.: Japan, Kyoto suburbs, vicinity of Kibuné Shrine.

Sabacon dentipalpe Suzuki, 1949: 8 View in CoL ; new ending.

dhaulagiri Martens, 1972: 314; loc. typ.: Nepal, Thakkola, ascend to Dapa-Col.

distinctus Suzuki, 1974a; loc. typ.: Japan, Kumamoto Pref., Kamimashiki-gun, Yabé-cho, Fusé-dô Cave.

Sabacon distinctum Suzuki, 1974a: 105 View in CoL ; new ending.

franzi Roewer, 1953a: 208 ; loc. typ.: Spain, Sierra de Ancares.

gonggashan Tsurusaki & Song, 1993: 158; loc. typ.: China, Sichuan Prov., Kangding, W slope of Mount Gong-ga-

shan.

imamurai Suzuki, 1964: 58; loc. typ.: Japan, Tokyo Pref., Nippara-dô Cave.

iriei Suzuki, 1974a: 103; loc. typ.: Japan, Kumamoto Pref., Yatsushiro-gun, Izumi-son, Ochiai, Yayama-daké-no-

tateana.

jiriensis Martens, 1972: 317; loc. typ.: Nepal, Jiri, ascend to Chordung.

ishizuchi Suzuki, 1974a: 100; loc. typ.: Japan, Ehime Pref., Mount Ishizuchi.

makinoi View in CoL makinoi Suzuki, 1949 View in CoL ; loc. typ.: Japan, Hokkaido, Nopporo.

Sabacon makinoi Suzuki, 1949: 15 View in CoL .

Sabacon habei Suzuki, 1965: 41 View in CoL ;—syn. Suzuki 1974a: 90.

Sabacon makinoi makinoi View in CoL — Suzuki & Tsurusaki 1983: 232.

makinoi View in CoL sugimotoi Suzuki & Tsurusaki, 1983: 232; loc. typ.: Japan, Akita Pref., L. Towada, Haka camping site.

martensi Tsurusaki & Song, 1993: 155 View in CoL ; loc. typ.: China, Sichuan Prov., Kangding, W slope of Mount Gong-ga-

shan.

mitchelli Crosby & Bishop, 1924 View in CoL ; loc. typ.: USA, North Carolina, Yancey Co., Mt. Mitchell.

Sabacon mitchelli Crosby & Bishop, 1924: 23 View in CoL ;— Shear 1975: 17 (redescr.).

occidentalis ( Banks, 1894b) View in CoL ; loc. typ.: USA, Washington, Olympia.

Phlegmacera occidentalis Banks, 1894b: 51 View in CoL .

Sabacon occidentalis View in CoL — Shear 1975: 19 (redescr.).

okadai Suzuki, 1941: 16; loc. typ.: China, Manchukuo, Moukden.

palpogranulatus Martens, 1972; loc. typ.: Nepal, Khumbu, Lughla.

Sabacon palpogranulatum Martens, 1972: 319 View in CoL ; new ending.

paradoxus Simon, 1879a View in CoL ; loc. typ.: France, Dép. Herault, caves.

Sabacon paradoxus Simon, 1879a: 266 View in CoL ;— Gruber 2003: 6; Prieto 2008: 51.

Sabacon paradoxum — Roewer 1914: 124; 1923: 695; Martens 1983: 274.

pasonianus Luque, 1991 View in CoL ; loc. typ.: Spain, Cantabria, Alto Asón-Miera.

Sabacon picosantrum View in CoL p-asonianum Luque, 1991: 18.

Sabacon picosantrum asonianum — Luque 1992: 144 (misspelling).

Sabacon picosantrum asoniarum — Luque 1992: 144 (misspelling).

Sabacon View in CoL p-asonianum — Luque 1992: errata.

Sabacon pasonianum View in CoL — Luque 1993: 131.

Sabacon pasonianus View in CoL — Prieto 2008: 51.

Remarks. The different spellings in Luque (1992) are listed in the errata of the same paper and probably reflect no flaw by the author.

picosantrum Martens, 1983: 292 View in CoL ; loc. typ.: Spain, Picos de Europa, Asturias, “Walking Stick Pot” Cave. Remarks. Gruber (in Prieto 2008: 51) considered the species epithet a noun in apposition and therefore did not alter the ending (IUCN 31.2.1).

pygmaeus Miyosi, 1942a; loc. typ.: Japan, Ehime Pref., Saraga-Miné Mountains.

Sabacon pygmaeum Miyosi, 1942a: 165 View in CoL ; new ending.

relictus Marten, 1972; loc. typ.: Nepal, Kathmandu Valley, Mount Phulchoki.

Sabacon relictum Marten, 1972: 320 ; new ending.

satoikioi Miyosi, 1942b View in CoL ; loc. typ.: Japan, Ehime Pref., Saraga-Miné Mountains.

Sabacon sato-ikioi Miyosi, 1942b: 49 View in CoL .

Sabacon satoikioi View in CoL — Suzuki 1974a: 100.

sergeidedicatus Martens, 1989; loc. typ.: Siberia, Altai Mountains, Cherga, Artybash, Teletskoye Lake .

Sabacon sergeidedicatum Martens, 1989: 372 ;— Chemeris & Logunov 2000: 210 (redescr.); new ending.

sheari Cokendolpher, 1984: 989; loc. typ.: USA, Oregon, Benton Co., Marys Peak.

simoni Dresco, 1952: 123 View in CoL ; loc. typ.: France, Dép. Alpes-Maritimes, Fôret de Turini.

siskiyou Shear, 1975: 22: loc. typ.: USA, California, Siskiyou Co., 3mi N McCloud.

unicornis Martens, 1972: 316; loc. typ.: Nepal, Gorapani Pass, betw. Pokhara and Kali-Gandaki Valley.

viscayanus View in CoL viscayanus Simon, 1881a View in CoL ; loc. typ.: Spain, Orduña, Cueva de la Embajada.

Sabacon viscayanus Simon, 1881a: 128 View in CoL .

Sabacon vizcayanus — Dresco 1952: 122; Gruber 2003: 6; Prieto 2008: 51.

Sabacon viscayanum viscayanum — Martens 1983: 277.

Sabacon viscayasanum viscayasanum — Luque 1992: 143.

Sabacon inflatus nomen nudum. Remarks. Dresco (1952: 118) found a specimen labelled inflatus in Simon’s collection to be identical with viscayanus View in CoL . The name reappears as a mere notice in Dresco (1955: 42) and Rambla (1967: 10).

viscayanus View in CoL ramblaianus Martens, 1983; loc. typ.: France, SW Lourdes, Val d’Ouzon, N. Ferrières.

Sabacon viscayanum ramblaianum Martens, 1983: 278 .

Sabacon viscayanus ramblaianus View in CoL — Prieto 2008: 51.

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Opiliones

Family

Sabaconidae

Loc

Sabaconidae Dresco, 1970

SCHÖNHOFER, AXEL L. 2013
2013
Loc

Sabacon altomontanus

Prieto, C. E. 2008: 51
2008
Loc

Sabacon pasonianus

Prieto, C. E. 2008: 51
2008
Loc

Sabacon viscayanus ramblaianus

Prieto, C. E. 2008: 51
2008
Loc

martensi

Tsurusaki, N. & Song, D. 1993: 155
1993
Loc

Sabacon pasonianum

Luque, C. G. 1993: 131
1993
Loc

Sabacon picosantrum asonianum

Luque, C. G. 1992: 144
1992
Loc

Sabacon picosantrum asoniarum

Luque, C. G. 1992: 144
1992
Loc

Sabacon viscayasanum viscayasanum

Luque, C. G. 1992: 143
1992
Loc

Sabacon picosantrum

Luque, C. G. 1991: 18
1991
Loc

crassipalpis

Martens, J. 1989: 370
Koch, L. 1879: 111
1989
Loc

Sabacon sergeidedicatum

Chemeris, A. N. & Logunov, D. V. 2000: 210
Martens, J. 1989: 372
1989
Loc

Sabacon bryanti

Shear, W. A. 1986: 5
1986
Loc

Sabacon altomontanum

Martens, J. 1983: 290
1983
Loc

Sabacon makinoi makinoi

Suzuki, S. & Tsurusaki, N. 1983: 232
1983
Loc

makinoi

Suzuki, S. & Tsurusaki, N. 1983: 232
1983
Loc

picosantrum

Prieto, C. E. 2008: 51
Martens, J. 1983: 292
1983
Loc

Sabacon viscayanum viscayanum

Martens, J. 1983: 277
1983
Loc

Sabacon viscayanum ramblaianum

Martens, J. 1983: 278
1983
Loc

Tomicomerus bryanti

Shear, W. A. 1975: 26
1975
Loc

Shear, W. A. 1975: 13
1975
Loc

Sabacon occidentalis

Shear, W. A. 1975: 19
1975
Loc

Sabacon akiyoshiense

Suzuki, S. 1974: 102
1974
Loc

Sabacon distinctum

Suzuki, S. 1974: 105
1974
Loc

Sabacon satoikioi

Suzuki, S. 1974: 100
1974
Loc

Sabacon palpogranulatum

Martens, J. 1972: 319
1972
Loc

Sabaconidae

Giribet, G. & Vogt, L. & Perez Gonzalez, A. & Sharma, P. & Kury, A. B. 2010: 421
Shear, W. A. 1986: 3
Martens, J. 1983: 269
Martens, J. 1978: 225
Dresco, E. 1970: 1210
1970
Loc

Hesperonemastoma inops

Gruber, J. 1970: 138
1970
Loc

Sabacon habei

Suzuki, S. 1974: 90
Suzuki, S. 1965: 41
1965
Loc

Sabacon akiyoshiensis

Suzuki, S. 1963: 156
1963
Loc

franzi

Roewer, C. F. 1953: 208
1953
Loc

simoni

Dresco, E. 1952: 123
1952
Loc

Sabacon vizcayanus

Prieto, C. E. 2008: 51
Gruber, J. 2003: 6
Dresco, E. 1952: 122
1952
Loc

Sabacon inflatus

Rambla, M. 1967: 10
Dresco, E. 1955: 42
Dresco, E. 1952: 118
1952
Loc

Sabacon dentipalpe

Suzuki, S. 1949: 8
1949
Loc

Sabacon makinoi

Suzuki, S. 1949: 15
1949
Loc

Sabacon jonesi

Shear, W. A. 1975: 8
Goodnight, C. J. & Goodnight, M. L. 1942: 1
1942
Loc

Sabacon pygmaeum

Miyosi, Y. 1942: 165
1942
Loc

Sabacon sato-ikioi

Miyosi, Y. 1942: 49
1942
Loc

Sabacon mitchelli

Shear, W. A. 1975: 17
Crosby, C. R. & Bishop, S. C. 1924: 23
1924
Loc

Tomicomerus bryantii

Roewer, C. F. 1914: 127
1914
Loc

Sabacon crassipalpis

Chemeris, A. N. & Logunov, D. V. 2000: 207
Martens, J. 1989: 370
Roewer, C. F. 1914: 125
1914
Loc

Sabacon paradoxum

Martens, J. 1983: 274
Roewer, C. F. 1923: 695
Roewer, C. F. 1914: 124
1914
Loc

Parasabacon Hansen & Sørensen, 1904: 83

Roewer, C. F. 1914: 123
Hansen, H. J. & Sorensen, W. 1904: 83
1904
Loc

Tomicomerus

Shear, W. A. 1986: 5
Pavesi, P. 1899: 533
1899
Loc

Tomicomerus bispinosus

Roewer, C. F. 1914: 127
Pavesi, P. 1899: 533
1899
Loc

Phlegmacera bryantii

Banks, N. 1898: 16
1898
Loc

Phlegmacera cavicoleus

Banks, N. 1901: 677
Banks, N. 1894: 52
1894
Loc

Phlegmacera occidentalis

Banks, N. 1894: 51
1894
Loc

Sabacon spinosus

Banks, N. 1894: 40
Weed, C. M. 1893: 575
1893
Loc

Phlegmacera

Roewer, C. F. 1914: 123
Packard, A. S. 1884: 203
1884
Loc

Phlegmacera cavicolens

Packard, A. S. 1884: 203
1884
Loc

Nemastoma inops

Shear, W. A. 2010: 107
Packard, A. S. 1884: 203
1884
Loc

Sabacon viscayanus

Simon, E. 1881: 128
1881
Loc

Sabacon

Simon, E. 1879: 266
1879
Loc

Sabacon paradoxus

Prieto, C. E. 2008: 51
Gruber, J. 2003: 6
Simon, E. 1879: 266
1879
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