Phanaeus (Notiophanaeus) halffterorum Edmonds, 1979

Kohlmann, Bert, Arriaga-Jimenez, Alfonsina & Roes, Matthias, 2018, Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae), ZooKeys 743, pp. 67-93 : 76-77

Phanaeus (Notiophanaeus) halffterorum Edmonds, 1979

Phanaeus halffterorum : Edmonds (1979: 99; partim), Halffter and Edmonds (1982: 88-89), Anduaga and Halffter (1991: 157), Delgado-Castillo et al. (1993: 125), Edmonds (1994: 39-43, 101), Anduaga (2000: 125, 130), Arnaud (2002: 95- 96), Edmonds (2003: 61, 65), Edmonds (2006: 31, 34, 36), Ceballos et al. (2009: 397), Edmonds and Zídek (2012: 3, 5, 12, 52, 54), Deloya et al. (2014: 77), Moctezuma and Halffter (2017: 52, 54-55), Moctezuma et al. (2017: 115), Lizardo et al. (in press).

Remarks.

This study identifies new localities of this species present in the Blackaller private collection, which seems to have a rather restricted distribution in the State of Mexico.

Estado de México (8 specimens): Sierra de Nanchititla, 6 km al Este de Nanchititla, Bosque de Pino-Encino, CD en Hongos, 1840 m, 15-VIII-2015, 18°52'N, 100°24'W, J.Blackaller y L. Zacarías cols.; 06-VIII-2011, Trampa de Luz, Blackaller y Robacker cols.; 21-VIII-2011, J. Blackaller col.

Comparing this species with its closest relative, Ph. bravoensis, one can add as a further difference between the two species the fact that Ph. halffterorum has its pygidium covered with coarse punctures while Ph. bravoensis has faintly impressed small punctures. Although it is not specified in the original description of this species (Edmonds 1979), specimens were collected directly by hand feeding on fungi pertaining to the genus Boletus.