Brachylinga laculata Webb, 2006

Webb, Donald W. & Metz, Mark A., 2006, A Revision of the New World Genera Brachylinga Irwin and Lyneborg and Lysilinga Irwin and Lyneborg (Diptera: Therevidae: Therevinae) with the Description of a New Genus, Elcaribe Webb, Zootaxa 1288 (1288), pp. 1-241 : 81-83

Brachylinga laculata Webb spec. nov.

( Figs. 14, 37, 60, 83, 106, 129, 152, 175, 195, 215)

Etymology

Laculatus (Latin) = four­cornered. This name is used as an adjective and refers to quadrate shape of the gonocoxite in lateral view ( Fig. 83).

Diagnosis

Brachylinga laculata is taxonomically associated with B. ornatifrons in possessing setae on the katepisternum; in having the halter knob dark reddish brown; and in having the foretibia dark yellow, apex dark reddish brown. Brachylinga laculata differs from B. ornatifrons in having the wing cell m 3 open widely; tergite 8 ( Fig. 14) thick medially; the gonocoxite ( Fig. 37) with a large gonocoxal bridge; and the distiphallus in dorsal view ( Fig. 129) tapered apically with a narrow dorsal carina.

Description of holotype male (MEI 076286)

Body length 6.0 mm.

Head. Length 0.64 mm. Ommatidia smaller ventrally and laterally. Frons pruinescence silver, black dorsally with small black circle dorsolateral to antennal base; setae black. Antenna/head length 0.67; black, pruinescence gray on scape; scape length 0.20 mm, width 0.14 mm, length/width 1.7, scape/pedicel length 2.7, scape/flagellum width 0.93, setae black, elongate, grading into macrosetae; pedicel length 0.07 mm, length/width 0.5; flagellum length 0.29 mm, width 0.15 mm, length/width 1.9, flagellum/ scape length 1.2. Parafacial pruinescence gray. Maxillary palpus dark brown, pruinescence gray, length 0.32 mm, length/width 5.3; setae pale brownish yellow. Genal setae white. Occipital setae white becoming pale brownish yellow dorsally. Postocular macrosetae dark reddish brown.

Thorax. Macrosetae dark reddish brown, 3 np, 2 sa, 1 pa, 1 dc, 2 sc. Mesonotum dark brown, pruinescence brownish gray; setae black, elongate intermixed with white setae laterally and across anterior margin. Pleuron and scutellum black, pruinescence gray; setae white on propleuron, anepisternum, katepisternum, laterotergite, metanepisternum, and scutellum. Wing. Membrane hyaline, veins brown; length 5.0 mm, length/width 2.8; pterostigma dark brown; cell m 3 open widely. Halter dark brown. Legs. Coxa setae white intermixed with dark brown setae, present on posterior surface of midcoxa; macrosetae dark reddish brown. Femora dark brown, pruinescence gray; ventral setae white intermixed with brown setae; av 0:0:5, pv 0:0:4. Tibiae dark yellow, apex reddish brown. Tarsomeres 1–2 dark yellow, apex reddish brown; 3–5 dark reddish brown.

Abdomen. Black, pruinescence white, dense; dorsal and lateral setae white, elongate. Terminalia (MEI 076250) dark brown, pruinescence white, dense. Tergite 8 ( Fig. 14), median length thick (0.09 mm); setae pale yellow across posterior margin. Sternite 8 ( Fig. 37), sides rounded, posterior margin emarginate; setae pale yellow. Epandrium ( Fig. 14), sides rounded; setae dark yellow. Cercus ( Fig. 14) ending above apex of hypoproct. Hypoproct ( Fig. 14), posterior margin emarginate, ending below apex of cercus. Gonocoxite ventral view ( Fig. 37) quadrate, medial margin slightly emarginate, tapering apically to short point; setae pale yellow with one dark brown setae apically; dorsal view ( Fig. 60) lacking posteromedial lobe, gonocoxal bridge large; lateral view ( Fig. 83) rectangular, posterior margin broadly pointed; ventral lobe ( Fig. 37) lacking ventral spines. Gonostylus ( Fig. 106), basodorsal lobe small; ventral margin curved, setae dark yellow, not produced apically in distinct tuft; fine, short spines absent laterally. Aedeagus with dorsal apodeme ( Fig. 129) narrow, quadrate, posterolateral margin slightly swollen; ventral apodeme ( Fig. 152) narrow, cylindrical, extending to anterior margin of dorsal apodeme; distiphallus dorsal view ( Fig. 129) tapered apically with narrow dorsal carina, lateral view ( Fig. 175) with apex curved ventrally at 90° angle, lacking apicolateral spines; ejaculatory apodeme ( Fig. 129), anterior margin truncate.

Variation (n=10). Body length 4.8–6.2, 5.5 mm.

Thorax. Macrosetae 3 np, 2 sa, 1 pa, 1 dc, 2 sc. Legs. Anteroventral macrosetae 0–1:0:2–4, pv 0:0:1–4.

Female

Similar to male except for following.

Variation (n=10).

Body length 5.9–7.2, 6.6 mm.

Head. Frons dark brown, pruinescence gray, whitish gray ventrally with small brown circle dorsolaterally; setae black, elongate. Antenna with setae on scape pale yellow intermixed with black setae.

Thorax. Macrosetae 3 np, 2 sa, 1 pa, 1 dc, 2 sc. Legs. Anteroventral macrosetae 0–1:0:3–4, pv 0:0:2–4.

Abdomen. Dark reddish brown, tergite 4 glossy, pruinescence gray on tergites 1 and 5 and posterolaterally on tergites 2–3; dorsal setae dark reddish brown and white, elongate, erect on tergite 1 and pruinescent areas of tergites 2–3. Terminalia (MEI 076179) dark reddish brown. Furca ( Fig. 195) rectangular, posterior margin rounded, sides parallel, anterior margin truncate; length 0.42 mm, width 0.26 mm; anterolateral projections absent. Common duct ( Fig. 195) shorter than furca. Spermathecal duct ( Fig. 195) diverging anteriorly.

Immature Stages. One female (MEI 040886) has been reared from a larva collected 30 km E La Ribera, Rancho Las Barracas from a sand habitat.

Distribution Brachylinga laculata is known from Mexico, Baja California Sur ( Fig. 215).

Habitats and phenology

Brachylinga laculata has been collected in pan traps with a few specimens collected in Malaise traps or handnetted on Prosopis juliflora (Sw.) DC. Adults have been collected from February to May and October and November.

Specimens examined

Type specimen. The holotype male of Brachylinga laculata Webb (MEI 076286) is labeled "Me., Baja Cal. Sur, Las Barracas, 17–IV 1985, Coll P. DeBach, pan trap" and is deposited in the CAS (Type No. 18114), on permanent loan from the UCR. PARATYPES: MEXICO. BAJA CALIFORNIA SUR. 11.3 km SE Guerrero Negro [27.921, ­114.102], 8.IV.1976, Doyen, J. T., Rude, P. A., 1♀ MEI 043542 ( EMEC) , Prosopis juliflora , 15 km S Damas de los Frailes [23.253, ­109.424] , 8.XI.1982, S.E., I.M., G.C.E., 2♀ MEI 081406, 081414 ( EMEC) ; 3.5 km WNW San Isidro [26.218, ­112.07] , 22–23.IV.1983, Wasbauer, M., 1♀ MEI 072515 ( CSCA) , Malaise trap; 30 km E La Ribera, Rancho Las Barracas [23.6, ­109.28] , 21–24.III.1982, MEI, EIS, 1♀ MEI 040886 ( MEIC) , sifted; 30 km E Santiago, Las Barracas [23.47, ­109.45] , 17–24.III.1982, DeBach, P., 1♀ MEI 082148 (E MEC); 25–31.III.1982, 2♀ MEI 081407, 081411 ( EMEC) , Malaise trap; 13–18.IV.1982, 2♀ MEI 040885, 082146 ( EMEC) ; 7–12.V.1982, 1♀ MEI 082070 ( EMEC) ; 8–14.X.1982, 1♀ MEI 082149 ( EMEC) ; 15–21.X.1982, 1♀ MEI 081404 ( EMEC) ; 22–28.X.1982, 2♀ MEI 081403, 082142 ( EMEC) ; 6–8.XI.1982, 1♀ MEI 081413 ( EMEC) ; 8–14.XI.1982, 3♀ MEI 081401, 081419–20 ( EMEC) ; 7–12.V.1982, 1♀ MEI 082145 ( EMEC) ; 6.4 km WSW Miraflores [23.347, ­109.817] , 22–23.IV.1979, Wasbauer, M., 19♂ MEI 072980–1, 072986, 072988–9, 072991–2, 072997–8, 073081, 073161, 073163, 073165, 073167, 073169, 073189–90, 073192, 073194, 30♀ 072630–2, 072979, 072982, 072985, 072987, 072990, 072993, 072994–6, 072999, 073064–5, 073067, 073075, 073160, 073162, 073170–6, 073188, 073191, 073193, 073195 ( CSCA) , Malaise trap; 23–24.IV.1979, 11♂ MEI 073061–4, 073179–83, 073187, 073202, 14♀ 073169, 073178, 073185–6, 073201, 073205–7, 073223–8 ( CSCA) , Malaise trap near water tank; Ciudad Constitucion [25.03, ­ 111.66], 22.II.1974, Bohart, G. E., 1♂ MEI 007716 ( EMUS) ; El Pescadero, Playa Los Cerritos [23.36, ­110.16], 14–15.IV.1979, Wasbauer, M., 1♀ MEI 073177 ( CSCA) , Malaise trap; Gulf of California , Isla Espirito Santo [24.478, ­110.344] , 23.III.1953, Arnaud, Jr., P. H., 3♂ MEI 042577, 042579, 042580 ( CAS) ; Gulf of California , Isla Partida [24.56, ­110.385] , 23.III.1953, 1♂ MEI 042578, 1♀ 042549 ( CAS) ; Las Barracas [23.472, ­109.453] , 11.X.1982, DeBach, P., 1♀ MEI 076138 ( UCR) ; 16.X.1982, 1♀ MEI 076136 ( UCR) ; 20–27.X.1982, 2♀ MEI 040700, 042520 ( UCR) ; 13.V.1983, 2♀ MEI 042532, 042534 ( UCR) ; 17.V.1983, 1♀ MEI 042539 ( UCR) ; 23.X.1983, 1♀ MEI 042536 ( UCR) ; 25.X.1983, 1♀ MEI 042526 ( UCR) ; 30.X.1983, 1♀ MEI 042531 ( UCR) ; 15.IV.1984, 1♀ MEI 042524 ( UCR) ; 16.IV.1984, 4♂ MEI 042527, 042529–30, 077506, 7♀ 042514–5, 042521–2, 076165, 076267, 076290 ( UCR) , yellow pan; 17.IV.1984, 2♂ MEI 076274, 076295, 2♀ 076164, 077500 ( UCR) ; 18.IV.1984, 4♀ MEI 042516, 042523, 076265, 076273 ( UCR) ; 19.IV.1984, 1♂ MEI 076256 ( UCR) ; 20.IV.1984, 1♂ MEI 076294, 3♀ ME I 040694, 042517, 076297 ( UCR) ; 21.IV.1984, 2♀ MEI 042537–8 ( UCR) ; 24.IV.1984, 1♂ MEI 076250, 2♀ MEI 042535, 042540 ( UCR) ; 28.IV.1984, 1♂ MEI 077505 ( UCR) ; 3.V.1984, 1♀ MEI 076179 ( UCR) ; 5.V.1984, 1♂ MEI 076190, 4♀ MEI 076183–5, 076271 ( UCR) ; 6.V.1984, 3♀ MEI 076186–7, 076278 ( UCR) ; 7.V.1984, 2♀ MEI 076188–9 ( UCR) ; 10.V.1984, 1♀ MEI 042518 ( UCR) ; 20.V.1984, 2♂ MEI 076252, 076255, 3♀ 076251, 076253, 076254 ( UCR) , pan trap; 26.V.1984, 2♂ MEI 042528, 076268, 4♀ 042519, 076167, 076176, 076281 ( UCR) ; 28.X.1984, 2♂ MEI 076102, 076133 ( UCR) ; 5.XII.1984, 1♀ MEI 076193 ( UCR) ; 16.XII.1984, 1♂ MEI 076260 ( UCR) ; 2.IV.1985, 3♂ MEI 076225, 076229, 076249, 13♀ 076216–23, 076226–8, 076230, 076235 ( UCR) ; 6.IV.1985, 1♂ MEI 076231, 3♀ 076232–4 ( UCR) ; 7.IV.1985, 7♂ MEI 076116–21, 076128, 15♀ 076110–5, 076122–7, 076129–31 ( UCR) ; 11.IV.1985, 7♀ MEI 076206–12 ( UCR) ; 17.IV.1985, 2♂ MEI 076169, 076171, 12♀ 076157, 076160–1, 076168, 076170, 076175, 076195–7, 076203–5 ( UCR) ; 21.IV.1985, 1♂ MEI 076141, 1♀ 076147 ( INHS) ; 23.IV.1985, 3♂ MEI 076259, 076296, 077507, 11♀ 076135, 076148, 076150, 076180, 076269, 076284–5, 076288–9, 076292, 076299 ( UCR) ; 27.IV.1985, 3♂ MEI 076174 (ME IC), 076177, 076262, 4♀ 076144 ( MEIC) , 076173, 076270, 076280 (UCR); 29.IV.1985, 3♀ MEI 076139, 076166, 076279 ( UCR) ; 30.IV.1985, 2♂ MEI 076257 ( CNC) , 076258 (USNM), 6♀ 076132 ( CNC) , 076172 (USNM), 076178, 076264, 077501, 077511 (UCR); 2.V.1985, 1♂ MEI 076215 ( UCR) ; 5.V.1985, 2♂ MEI 076198, 076202, 4♀ MEI 076162, 076199, 076200–1 ( UCR) ; 11.V.1985, 1♂ MEI 077502 ( INHS) , 4♀ MEI 076194 ( INHS) , 076277, 077504, 077508 (UCR); 13.V.1985, 1♀ MEI 077503 ( UCR) ; 14.V.1985, 1♀ MEI 077510 ( UCR) ; 17.V.1985, 1♂ MEI 040696 ( UCR) ; 20.V.1985, 4♀ MEI 076143, 076149, 076151–2 ( UCR) ; 27.V.1985, 2♂ MEI 076145, 076154, 7♀ 076137, 076140, 076156, 076181–3, 076291 ( UCR) ; 27.V.1985, 1♀ MEI 076293 ( UCR) ; 1.VI.1985, 1♂ MEI 077509, 2♀ 076134, 076142 ( UCR) ; 21.X.1985, 2♂ MEI 076103, 076191, 2♀ 076266, 076275 ( UCR) ; 5.XI.1985, 2♀ MEI 076192, 076282 ( UCR) ; 11.IV.1986, 1♀ 076272 ( UCR) ; 21.IV.1986, 1♀ MEI 076287 ( UCR) ; 2.V.1986, 2♂ MEI 076261, 076263, 1♀ 076153 ( UCR) ; 2.V.1986, 1♀ MEI 076276 ( UCR) ; 10.V.1986, 2♂ MEI 076243, 076248, 5♀ 076241, 076244–7 ( UCR) ; 16.V.1986, 2♂ MEI 076237–8 3♀ 076239–40, 076242 ( UCR) ; 26.V.1986, 1♂ MEI 076213, 2♀ 076214, 076236 ( UCR) ; 15.IV.1987, 1♂ MEI 076155 ( UCR) ; Los Barriles [23.69, ­109.698], 27–28.IV.1979, Wasbauer, M., 3♀ MEI 073196, 073199–200 ( CSCA) , Malaise trap; Los Animas , Sierra Lagun [23.567 ­110.000] , 12.X.1941, Ross, Bohart, 1♂ MEI 043230 ( CAS) .