Hierochloe redolens var. gunckelii ( Parodi 1941: 197 ) Lema, Sahuquillo & Pimentel, 2021

Lema-Suárez, Irene, Sahuquillo, Elvira & Pimentel, Manuel, 2021, A revision of Hierochloe sect. Monoecia (Anthoxanthinae, Pooideae, Poaceae), Phytotaxa 478 (1), pp. 92-104 : 99

4.2. Hierochloe redolens var. gunckelii ( Parodi 1941: 197) Lema, Sahuquillo & Pimentel comb. nov.

Basionym: Hierochloe gunckelii Parodi (1941: 197) .

Type: CHILE. [Cultivated in the Jardín Botánico de la Facultad de Agronomía de Buenos Aires] Valdivia Province , Corral (Amargos), Sept. 1940, L.H. Gunckell s.n. (holotype: BAA-00000183 digital image!; isotypes: BAA-00000296 digital image!, Kn.v., SI n.v.) .

Homotypic names: Anthoxanthum gunckelii (Parodi) Veldkamp in Schouten & Veldkamp (1985: 348). Anthoxanthum redolens (Vahl) P. Royen var. gunckelii (Parodi) N. Villalobos & Finot (2019: 124) .

Description: Robust plant, (50) 100–150 cm. Ligules obtuse. Leaf-blades (15)25–35(50) cm × 5–7(11) mm, convolute, scabrous adaxially and glabrous, scaberulous or scabrous abaxially. Panicle 10–25 × 1.5–3 cm, open or slightly contracted, lanceolate, equilateral or nodding. Lower branches 5–7 cm long, geminate. Panicle branches pubescent, without spikelets in the lower third. Spikelets 5.5–7 mm, cuneate. Glumes ovate, 3-veined. Lower glume 4–6 mm; upper glume 5–7 mm. Lemma of male florets 5–7 mm, elliptic or oblong, papyraceous. Midvein and margins of the lemma scabrous, eciliate or ciliolate. Awn of lower male floret 0.3–1.5 mm, subterminally inserted. Awn of upper male floret 1.5–2 mm, straight, dorsally inserted. Lemma of apical female floret 4.3–4.8 mm, membranaceous, scabrous. Midvein eciliate or ciliolate.

Leaf anatomy: Long cells 180–250 μm. Stomata in adaxial surface. Hooks present but scarce. Bulliform cells absent. Leaf section open with 30–45 quadrangular ribs ( Figure 1. D ).

Habitat: Sandy or dry soils in coastal areas ( De Paula 1975, Sede 2012).

Distribution: Grows in Central-South Chile (from Biobio to Los Lagos, including Chiloe Island) and Central Argentina (Rio Negro) ( De Paula 1975, Sede 2012).

Genome size and DNA-ploidy levels: 2C = 19.06–19.64 pg, 6 x.

Phenology: September–January.

Specimens examined:— CHILE. Los Lagos Region: Chiloe Province: Isla de Chiloe, Caleta Compu, next to church. Meadow near the sea, behind a formation dominated by Puccinellia glaucescens and Salicornia sp. , 13 m, S 42º 52.300’ W 073º 42.086’, 17 January 2014, E. Sahuquillo & M. Pimentel s.n. (SANT-73538).

Figures

FIGURE 1. Hierochloe sect. Monoecia micromorphology details.A–C. Leaf transverse section. A. H. pusilla, leaf transverse section open with round ribs. B. H. juncifolia, leaf transverse section closed with polygonal ribs. C. H. redolens var. altissima, leaf transverse section open with quadrangular ribs. D–F. Leaf epidermis. D. H. redolens var. gunckelii, adaxial epidermis with long cells type I3 and stomata. E. H. pusilla, leaf epidermis with long cells type I3 and short cells S5 (round). F. H. juncifolia, leaf epidermis with long cells type I3 and short cells S2 (ax shaped).