Montenegrina perstriata drimica Nordsieck, 1972

Feher, Zoltan & Szekeres, Miklos, 2016, Taxonomic revision of the rock-dwelling door snail genus Montenegrina Boettger, 1877 (Mollusca, Gastropoda, Clausiliidae), ZooKeys 599, pp. 1-137 : 64-66

Taxon classification Animalia Stylommatophora Clausiliidae

Montenegrina perstriata drimica Nordsieck, 1972 Fig. 27D, E

Montenegrina perstriata drimica Nordsieck, 1972: 32, plate 4, fig. 38. - Zilch 1981: 130, plate 14, fig. 31. - Nordsieck 2009: 74, plate 1, fig. 3. - Dedov and Neubert 2009:91, plate 1, figs 3-4.

Montenegrina perstriata crassa Erőss & Szekeres, 1999 in Erőss et al. 1999: 448, fig. 3. - Nordsieck 2009: 75.

Diagnosis.

Shell small to large, tumid, light to dark corneous. Lower whorls smooth, flattened, nearly of the same width. Upper whorls finely striate-costate. Neck densely striate-costate. Peristome with weakly to very strongly swollen, reflexed margin. Lamellae superior and spiralis overlap. In front view lamella subcolumellaris often visible. Lunella lateral, separate from the weak basalis. Subclaustralis short, sulcalis present. Anterior plica superior absent or weak, separate from the lunella complex. Clausilium plate not visible through the aperture.

Dimensions

(in mm). Hs: 14.0-21.5 (holotype 16.5), Ws: 3.8-5.0 (holotype 4.3).

Type locality.

Macedonia, valley of the Crni Drin, Lukovo near Struga [ drimica ]; Macedonia, valley of the Crni Drin, left side, 2 km N of Lukovo and 1 km S of the road junction to Modrić, 41.3654°N, 20.6048°E [crassa].

Type material.

Lukovo near Struga, leg. H. Nordsieck, 27.viii.1971, holotype [ drimica ] (SMF 221048), paratypes [ drimica ] (SMF 221049, SMNS-N 5500/30); same locality, leg. H. Nordsieck, 19.viii.1971, paratype [ drimica ] (SMNS-N5596); 2 km N of Lukovo, leg. ZE, ZF, 13.viii.1996, holotype [crassa] (HNHM 70834), paratypes [crassa] (HNHM 70835/6, NHMW 100076/2, SMF 312509/2, NMBE 535059/1, NMBE 22592/3).

Other material.

Macedonia, Debar District, S of Debar, at the dam of the Crni Drin, 590 m, 41.4948°N, 20.5052°E, leg. ZF, EH, KJ, HS, 14.x.2014 (NHMW 110430/MN/0087); between Lukovo and Džepište, leg. Sipos, AS, Topál, 13.x.1975 (HNHM 10139/35); 1 km SE of Džepište, leg. LP, PS, AS, 17.vii.1972 (HNHM 36712); Struga District, valley of the Crni Drin, 3.7 km S of Lukovo, 700 m, 41.3299°N, 20.6369°E, leg. ZF, EH, KJ, HS, 15.x.2014 (NHMW 110430/MN/0088); same locality, leg. ZE, ZF, AH, 8.iv.2004 (HNHM 94448); 3 km S of Lukovo, Brana Globočica, 41.3355°N, 20.6345°E, leg. ZE, ZF, AH, 8.iv.2004 (HNHM 94447); 2 km S of Lukovo, S of the viaduct, 690 m, 41.3397°N, 20.6273°E, leg. ZF, EH, KJ, HS, 15.x.2014 (NHMW 110430/MN/0089); same locality, N of the viaduct, 690 m, 41.3406°N, 20.6266°E, leg. ZF, EH, KJ, HS, 15.x.2014 (NHMW 110430/MN/0090); same locality, leg. ZE, ZF, AH, 8.iv.2004 (HNHM 94446); Lukovo, S edge of the village, 640 m, 41.3545°N, 20.6124°E, leg. ZF, EH, KJ, HS, 15.x.2014 (NHMW 110430/MN/0091); same locality, leg. ZE, ZF, AH, 8.iv.2004 (HNHM 94445); 1 km N of Lukovo, 630 m, 41.3623°N, 20.6036°E, leg. ZF, EH, KJ, HS, 15.x.2014 (NHMW 110430/MN/0092); 1.3 km N of Lukovo, 620 m, 41.3638°N, 20.6049°E, leg. ZF, EH, KJ, HS, 15.x.2014 (NHMW 110430/MN/0093); 1.5 km N of Lukovo, 620 m, 41.3666°N, 20.6049°E, leg. ZF, EH, KJ, HS, 15.x.2014 (NHMW 110430/MN/0094); same locality, leg. ZE, ZF, AH, 8.iv.2004 (HNHM 94449); 1.7 km N of Lukovo, 620 m, 41.3672°N, 20.6046°E, leg. ZF, EH, KJ, HS, 15.x.2014 (NHMW 110430/MN/0095); 2 km N of Lukovo, 620 m, 41.3695°N, 20.6021°E, leg. ZF, EH, KJ, HS, 15.x.2014 (NHMW 110430/MN/0096); same locality, leg. ZE, ZF, AH, 8.iv.2004 (HNHM 94589); 2.4 km N of Lukovo, S of the Modrić junction, 610 m, 41.3705°N, 20.5979°E, leg. ZF, EH, KJ, HS, 15.x.2014 (NHMW 110430/MN/0097); 2.5 km N of Lukovo, at the Modrić junction, 610 m, 41.3713°N, 20.5971°E, leg. ZF, EH, KJ, HS, 15.x.2014 (NHMW 110430/MN/0098); 6 km N of Lukovo, side road to Lokov, 620 m, 41.3978°N, 20.5899°E, leg. ZF, EH, KJ, HS, 15.x.2014 (NHMW 110430/MN/0099); 4 km N of Lukovo, 600 m, 41.3816°N, 20.5881°E, leg. ZF, EH, KJ, HS, 15.x.2014 (NHMW 110430/MN/0100).

Distribution.

Upper valley of the Crni Drin between the Globočica Lake and Debar (Fig. 25).

Remarks.

This taxon shows considerable intra- and inter-population heterogeneity in terms of shell size and peristome thickness. Nordsieck (2009) questioned the distinct subspecific status of crassa, which he regarded "a local form of Montenegrina perstriata drimica with thickened peristome, anterior upper palatal plica continuous with upper palatal plica …”. He argued that "there are further forms of Montenegrina perstriata drimica with thickened peristome occurring farther in the south" ( Nordsieck 1972: 32-33). Based on these arguments, Dedov and Neubert (2009) already treated crassa as the junior synonym of drimica , and we also accept this view.