Eukraiohelea inusitata, Johannsen, 1946

Johannse, O. A., 1946, Some New Species Of Nemocerous Diptera From Guam, Insects of Guam II, Honolulu, Hawaii: Bernice P. Bishop Museum, pp. 187-193 : 190-191

5. Eukraiohelea inusitata , new species (fig. 1, i).

Female: head and large basal segment of antenna yellowish brown; palpi, mouth parts, and flagellum of antenna, dark brown. Antenna short bristly haired, slender, about two thirds as long as the body, intermediate segments fusiform, terminal segments slender, cylindrical, progressively increasing in length. Thorax dusky yellow, ground color of mesonotum brown, slightly shining, with three slender pruinose vittae that are transversely connected beyond the middle by some broader pruinose bands; laterad of the vittae are several pruinose spots. Posterior margin of the scutellum pruinose. Abdomen brown, posterior margins of the segments pruinose, more extended on posterior segments. Legs, including coxae, yellow, tips of tibiae blackened, fourth tarsal segment short and strongly bilobed, fifth segment longer and more slender, each with one long claw and a short basal lateral tooth; hind tibiae broadened at tip. Wing venation as figured (fig. 1, i) costa bristly; petiole of radius with a pair of short bristles before the middle; other veins bare, the wing margin fringed; microtrichia present, macrotrichia absent. The wing spots that narrowly margin the tips of the veins, and more prominently the crossvein, and at the angle of the radial sector near its tip and a small crescent beyond this, owe their brown color to coarser microtrichia. Wing surface faintly yellow tinged, costa and radial veins yellow. Halteres dark brown. Fore femora each with a single spine near the middle on the under side. Length 2.2 mm.; wing, measured from the humeral crossvein, 1.9 mm.

Agana , June 26, Usinger.

Holotype in the Cornell University collection.

Ingram and Madie have described two closely related species, the first as Parabezzia poikiloptera (Ann. Trop. Med. 16: 278, 1922), the second as Stilobezzia ugandae (Bull. Ent. Res. 14: 62, 1923). The wing venation in both is similar to the above described species, but they differ in lacking the spine on the underside of the fore femur. By definition, the Guam species falls in with the genus Eukraiohelea of Ingram and Macfie (Ann. Trop. Med. 15:347, 1921), because of the presence of the femoral spine; but in other characters it seems more closely related to P. poik-iloptera. Most species of Stilobezzia have two anterior radial cells, whereas the three species noted above have but one. Whether it is more expedient to include these and other similar forms in Stilobezzia or whether they should be placed in Eukraiohelea by broadening its scope is left for the future. It seems that the most reasonable course would be to broaden the scope of Stilobezzia and include the aberrant forms under subgenera. Until the female of Parabezzia petiolata Malloch is found, the position of the genus Parabezzia with respect to related genera is in doubt.