Hypharpax australis ( Dejean, 1829 )

Kataev, Boris M., 2021, On some Afrotropical Harpalus, with description of two new species, and remarks on Hypharpax australis (Coleoptera: Carabidae): misidentification, mislabeling and introduction to the Australian region, Zootaxa 5020 (1), pp. 31-56 : 49

Hypharpax australis ( Dejean, 1829)

( Figs 45, 46 )

Harpalus australis Dejean, 1829: 385 .

Harpalus inornatus Germar, 1848: 169 .

Harpalus coxii Laporte de Castelnau, 1867: 107 .

Hypharpax abstrusus Bates, 1878: 23 .

Hypharpax parvus Chaudoir, 1878: 500 .

Type material examined. Holotype of Harpalus australis : ♂, labeled “ ♂ ”, “australis m in Nov. Holland ” [ Dejean’s handwriting], “TYPE”, and “ Hypharpax australis Dej. examined. 26.IX.78” ( MNHN).

Lectotype of Hypharpax abstrusus [designated by Larochelle & Larivière (2005)]: ♂, “ Auckland N. Z.”, “Hy- pharpax abstrusus Bates ” [Bates’ handwriting], “Ex Musaeo H.W. BATES”, “ LECTOTYPE ♂ Hypharpax abstrusus Bates, 1874 designated by Larochelle & Larivière, 2004” ( MNHN).

Syntypes of Hypharpax parvus : 2 ♂♂, 1 ♀, with a common bottom label “ parvus Chaudoir Australie ” [Chau- doir’s handwriting] ( MNHN) .

Additional material examined. Australia: 1 ♂, “Parkwille Vic., N.O. 36, A. Wade”, “ Hypharpax australis Dej., Id. by R. Blackwood”, “ Hypharpax australis ( Dejean, 1829) , Det. A. Larochelle 2004” ( ANIC) ; 1 ♀, “ Blue Mts. , N.S. Wales ”, “E.W. Ferguson Collection”, “ Hypharpax australis Dej., N.S.W. ”, “ Hypharpax australis ( Dejean, 1829) , Det. A. Larochelle 2004” ( ANIC) ; 1 ♂, “ Melbourne, No. 19, Ejnar Fischer ”, “19”, “ Diaphoromerus austral- asiae Dej., Id. by T.G. Sloane ”, “ Harpalus australasiae Dejean, 1829 , Det. A. Larochelle 2004” ( ANIC) ; 1 ♂, “Mel- bourne, No. 19, Ejnar Fischer”, “ Hypharpax australis Dej., Id. by T.G. Sloane ” ( ZMUN) ; 1 ♀, “ Austral ”, “CNHM, 1955, Karl Brancsik Colln., Ex Eduard Knirsch”, “ Notiobia (Anisotarsus) australasiae (Dejean) Det. G.R. Noonan 1973 ” ( FMNH) ; 1 ♀, with round piece of reddish paper, and “Nova Hollandia. H. australis Dej. ” ( ZSM) ; 1 ♀, “ Nov. holl.”, “ australis Dej. Nov. holl. [Ménétriés’ handwriting]” ( ZIN) ; 1 ♂, Canberra , vall. Jerrabomberra R., marshes, on dump soil, 7.XI.1999, I. Sokolov leg. ( ZIN) . Tasmania: 1 ♂, Launceston, Cataract Gorge Reserve , 9–12.XI.1999, V. Zherikhin leg. ( ZIN) ; 1 ♀, “ Launceston ”, “Ex. Coll. S.A. Mus. ”, “ Hypharpax australis ( Dejean, 1829) , Det. A. Larochelle 2004” ( ANIC) .

New Zealand: 1 ♀, “Cristchurch, Maffeys Road, 23.IX.1995 R.M. Emberson, running on ground”, “ Hypharpax australis (Dejean) , Det. R.M. Emberson 1996” ( ZIN) ; 1 ♂, “Cristchurch, MC Port Hills , in garden, 13.IV.1985 P. Syrett. ”, “ Hypharpax australis (Dejean) , Det. R.M. Emberson 1996” ( ZIN) .

Distribution. Widely distributed in Australia, including Tasmania and Lord Howe Island (Moore in Moore et al. 1987); introduced to New Zealand where it is now also widely distributed ( Larochelle & Larivière 2005).

Comparison. As a member of Anisodactylina , this species distinctly differs from Harpalus parvulus and H. australasiae in having paragossae glabrous, male pro- and mesotarsi with spongy adhesive vestiture ventrally, and aedeagus with apical capitulum in dorsal position. In external features, it is recognizable by relatively wider head and pronotum, and additionally differs from H. parvulus in having pronotum much more strongly narrowed basally and with basal angles more widely rounded, and from H. australasiae in having metafemur ventrally with four or five setae along posterior margin and pronotal basal edge with distinct setae. The dorsal microsculpture in Hypharpax australis is very distinct, throughout clearly isodiametric.

Figures

FIGURES 45, 46. Hypharpax australis (Dejean, 1829) (45, New Zealand; 46, holotype), habitus. Scale bar = 1.0 mm.