Tonnoiriella arcuata Ježek, 1997

Wagner, Rüdiger & Withers, Phil, 2020, The West-Palearctic species of the genus Tonnoiriella Vaillant, 1971 (Diptera: Psychodidae, Psychodinae), Zootaxa 4728 (2), pp. 183-210 : 205

Tonnoiriella arcuata Ježek, 1997

( Figs 48, 49 )

Tonnoiriella arcuata Ježek, 1997 : Časopis Národniho Muzea v Praze Řada přirodovedna, 166: 1 16.

Material: 1♂ holotype, slide mounted: head, mothparts with 2 palpi, 2 antennae, thorax cut in half, 2 wings, legs, abdomen part, 1 stylus, aedeagus, epandrium with epandrial processes; left side white label: Caucasus, Pskhu, 27.6.1988 Ježek leg.; bowl shaped basin (village Pskhu). Swamp slopes nr. Stream Betaga, 2.000 m a.s.l., mountain forest, Alnus , Corylus , Sambucus , Fragaria, Musci Hepaticae. Right side red label: Tonnoirella arcuata Ježek 1999 , holotype Cat. No. 33663. Paratypes: Cat. Nos. 33666, 33667, 33669; collected with holotype (all in Natural History Museum Prague).

Description: Head with eyebridge of 3 facet rows; distance between eyes 5.5 facet diameters; one row of postocular bristles. Antenna with scape, pedicel and 14 flagellomeres: absolute length 0.042-0.042-0.036-0.036-0.036- 0.036-0.034-0.034-0.03-0.028-0.022-0.026-0.026-0.018-0.02- 0.03 mm; relative length 21-21-18-18-18-18-17- 17- 15-14-11 -13-13-9-10-15; terminal flagellomere with apiculus. Pairs of digitate ascoids on flagellomeres 5–11; one antenna with two flagellomeres fused so that the flagellum appears 13-segmented. Palpus segments absolute length: 0.048-0.068- 0.084-0.134 mm; relative length 24-34-42-67.

Wing 2.95 times longer than wide; length 1.67 mm. width 0.57 mm.

Male terminalia with thin hypandrium with 2 small lateral angular projections. Gonocoxites (probably artificially compressed) are approximately 2x longer than wide, with a long seta in the distal fourth, condyles pentagonal. Gonostyli 1.3 times longer than gonocoxites, slightly bent. Parameres small, not clearly discernable.

Aedeagus ( Fig. 48 ) with short basiphallus sclerite; ventral arm of basiphallus bilobed, laterally connected to a U-shaped ventral distiphallus sclerite. Dorsal lobe of basiphallus broad straight, no distiphallus sclerite attached.

Epandrium ( Fig. 49 ) not clearly detectable; epandrial processes elongate, almost straight with 8 tenacula, apical ones shorter than basal ones.

Distribution: Caucasus mountains.

Remarks: A species, easy to identify by the the double-headed hypandrium, and the distiphallus sclerite, almost semicircular and evenly bent.

Figures

FIGURES 43–49: 43–45 T. nigricauda 43 hypandrium, gonopods, aedeagus, 44 aedeagus ventral view; 45 variability of the ventral distiphallus sclerite in England, Poland, North- and South Germany (left to right?); 46 T. goncalvesi genitalia ventral view; 47 T. andradei hypandrium, gonopods, aedeagus; 48–49 T. arcuata (holotype) 48 hypandrium, gonopods, aedeagus, 49 epandrium and epandrial processes, oblique view.