Tonnoiriella atlantica ( Satchell, 1955 )

Wagner, Rüdiger & Withers, Phil, 2020, The West-Palearctic species of the genus Tonnoiriella Vaillant, 1971 (Diptera: Psychodidae, Psychodinae), Zootaxa 4728 (2), pp. 183-210 : 197-198

Tonnoiriella atlantica ( Satchell, 1955)

( Figs. 26–28 )

Pericoma atlantica Satchell, 1955 : Annals of the Natal Museum 13: 112.

Tonnoiriella atlantica ; Wagner 1987: Entomofauna 8 (2): 14.

Material: 1♂ holotype on pin: 4 microslides with head, styles + aedeagus, wings, thorax and abdomen, red la- bel: (handwritten) Pericoma atlantica Satchell m.s., pink label: Peric (Lib) atlantica holotype, with label with red margin: type ♂; circular white label: El Biar 1.II. [18]93, reverse Lidad atlantica; white label: Eaton Bequest B.M. 1929-590. [Note: El Biar is a district of Algiers, the capital of Algeria].

1♂ slide mounted; round white label (partly pasted over): palpus, antenna, hind leg, not readable, penis; white label Eaton no2 c; white label: Eaton Bequest 1929-590; white label: Tonnoirella atlantica ; blue label: paratype Pericoma atlantica Satchell m.s.; white label: Libidophila atlantica ♂, Torrent beyond l’Hospice de Vieilliards, Bône, Algeria, 14.III.1903 Alg. No. VI. (partly pasted over by small blue label: paratype (all BMNH). The slide contains 2 wings, 1 leg, 2 antennae, 1 palpus complete, 1 palpus incomplete, epandrial processes lateral view, epandrium and epandrial processes lateral view, 2x gonocoxite and gonostylus, 1x aedeagus lateral view.

Further material: 1♂ 1♀, Tunisia: Oued Titria , 5 km östlich Ain Sobah, 90 m a.s.l., 36°56’N / 8°56’E, 3 June 1982, shaded stream ; 2♂ 2♀ Tunisia: 4 km südlich Ain Drahan , 530 m a.s.l., 36°43’N / 8°40’E, 17.-18. May 1982, stream; all leg. Malicky ( RW) .

Description (holotype): head missing; antenna of scape pedicel and 14 flagellomeres. Relative length of antennomeres: 21-26-24-24-26-26-28-28-27-24-20-19-17-14-13-17; absolute length: 0.04-0.05-0.05-0.05-0.05-0.05- 0.06-0.06-0.05-0.05-0.04-0.04-0.03-0.03-0.03- 0.03 mm; flagellomeres 5–11 with a pair of digitate ascoids. Palpus four segmented; relative length: 38-41-60-71; absolute length: 0.08-0.08-0.12- 0.14 mm.

Thorax missing. Wing length 2.17 mm, width 0.82 mm; length by width relation: 2.66.

Male terminalia with gonocoxites approximately 3x longer than wide, parameres not discernable; gonostyli almost straight, bent only near tip. Epandrial processes ( Fig. 28 ) in lateral view 2x longer than high, with 5–6 feathered tenacula, 3 almost as long as epandrial processes, 2 about half as long.

Description of a T. atlantica specimen from Tunisia: Head with eyebridge of 3 facet rows; distance between eyes 3 facet diameters; with postocular bristles. Antenna with scape, pedicel and 14 flagellomeres; absolute length of antennomeres: 0.040-0.052-0.040-0.040-0.040-0.042-0.042-0.042-0.040-0.036-0.036-0.034-0.034-0.028- 0.026- 0.032 mm; relative length: 20-26-20-20-20-21-21-21-20-18-18-17-17-14-13-16. Flagellomeres 5–11 with a pair of digitiform ascoids. Absolute length of palpus segments: 0.060-0.088- 0.106-0.186 mm; relative length: 30-44-53- 93.

Wing length by width relation 2.61; length 1.86 mm, width 0.71.

Male terminalia with a thin hypandrium with linguiform median extension ( Fig. 26 ). Gonocoxite about 3x longer than wide with a long seta at about middle on inner surface. Gonostylus little longer than the gonocoxite, slightly bent.

Parameres with well-developed ventral bridge (pocket shaped oval lobes), median portion not discernable; dorsal parts large inner edges bent.

Aedeagus with basiphallus as long as gonocoxite with darker sclerotization along the mid-line. Ventral lobe curved bipartite; ventral arm connected with the cross distiphallus dorsal arm almost straight; the dorsal lobe is very faint, straight and is distally with a faint sclerite. Distiphallus with a triangular basis connected to the hook shaped ventral basiphallus lobe that is slightly bent apically.

Epandrium rhomboid with 2 foramina, epandrial processes with 6 tenacula, 3 long and 3 short on one side, 2 long and 4 short on the other side ( Fig. 27 ).

Distribution: Algeria, Tunisia.

Remarks: Type specimens from Algeria and specimens from Tunisia obviously belong to the same species, based on the shape of the hypandrium and the aedeagus sclerites.

Figures

FIGURES 26–31: 26–28 T. atlantica: 26 hypandrium, gonopods, aedeagus; 27 epandrium, epandrial processes, proctiger; 28 epandrial processes with retinacula, lateral view; 29–31 T. fontinalis: hypandrium, gonopods, aedeagus of specimen from Corsica (29), Sardinia (30), parameres and condyles of specimens from Corsica (31).