Pupina crosseana Morlet, 1883

Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak, 2022, Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia, ZooKeys 1119, pp. 1-115 : 1

Pupina crosseana Morlet, 1883

Figs 28H-J , 30E, F , 31A

Pupina crosseana Morlet, 1883: 108, 109, pl. 4, fig. 5. Type locality: Cambodge [Cambodia]. Morlet 1889: 152, Pnom-Rohan (Cambodge) [Phnum Roung, Kampong Thom Province, Cambodia]; Ajuthia (Siam) [Phra Nakhon Si Ayutthaya Province, Thailand]. Fischer 1891: 107. Morlet 1904: 371, pl. 20, fig. 14, 14a. Fischer and Dautzenberg 1904: 431. Fischer-Piette 1950: 153. Fischer 1973: 48. BEDO 2017: 89.

Pupina (Tylotoechus) crosseana - Kobelt 1902: 310, 311. Hemmen and Hemmen 2001: 39.

Type material examined.

Lectotype MNHN-IM-2000-35834 (Figs 28H , 30E ) from Cambodge. Paralectotype RBINS MT966/10591 (1 shell; Figs 28I , 30F ) from Phnom-Rohan, Cambodge.

Other material examined.

CUMZ 12049 (16 shells; Figs 28J , 31A ) from Khao Jedee Temple, Ta Kli District, Nakhon Sawan Province, 25 Oct. 2005.

Diagnosis.

Shell fusiform; last whorl ca. three quarters of shell height. Apertural lip somewhat thickened, but not expanded; apertural lip when observed from lateral view somewhat curved. Columellar tooth fin-shaped.

Differential diagnosis.

Pupina crosseana is most similar to P. perakensis in having a fusiform shell shape, but differs in having the parietal callus and parietal tooth less thickened, a less curved apertural lip when observed from lateral view, and a fin-shaped columellar tooth.

Distribution.

Cambodia and central Thailand ( Fischer and Dautzenberg 1904).

Remarks.

As the original description did not explicitly state that the description of this species was based on a single specimen (nor could this be inferred), the designation of a holotype by Fischer-Piette (1950) in fact constitutes a lectotype designation ( ICZN 1999: Art. 74.6).

Figures

Figure 28. Shells of Pupina arula species group from mainland Southeast Asia A-G Pupina peguensis A syntype of Pupina blanfordi NHMUK 1888.12.4.100 and specimens B CUMZ 12105 C CUMZ 12050 D CUMZ 12103 E NHMUK ex. Cuming coll. F CUMZ 12094, and G CUMZ 12108 H-J Pupina crosseana H lectotype MNHN-IM- 2000 - 35834 I paralectotype RBINS MT 966 / 10591, and J specimen CUMZ 12049 K, L Pupina siamensis K lectotype SMF 109948 and L specimen CUMZ 12052 M-P Pupina bilabiata sp. nov. M holotype CUMZ 12073 / 1 and specimens N CUMZ 12079 O CUMZ 12081, and P CUMZ 12086 Q, R Pupina godwinausteni sp. nov. Q holotype CUMZ 12090 / 1 and R paratype CUMZ 12090 / 2. Photo: P. Maestrati, MNHN (H), F. Trus, RBINS (I).

Figure 30. A-D Pupina peguensis: specimens A CUMZ 12050 from Chai Thong Wararam Temple, Nakhon Sawan B CUMZ 12105 from Thep Phithak Punnaram Temple, Nakhon Ratchasima C CUMZ 12108 from Tham Thep Bandan Temple, Phetchabun, and D CUMZ 12094 from Khao Tham Phra Temple, Chiang Rai E, F Pupina crosseana E lectotype MNHN-IM- 2000 - 35834 from Cambodge and F paralectotype RBINS MT 966 / 10591 from Phnom-Rohan, Cambodge. Photo: P. Maestrati, MNHN (E), F. Trus, RBINS (F).

Figure 31. A Pupina crosseana, specimen CUMZ 12049 from Khao Jedee Temple, Nakhon Sawan B, C Pupina siamensis: B lectotype SMF 109948 and C specimen CUMZ 12052 from Sri Thong Cave, Sra Keo D-F Pupina bilabiata sp. nov. D holotype CUMZ 12073 / 1, and specimens E CUMZ 12079 from Tham Pha Ya Temple, Loei and F CUMZ 12081 from Pha Jor Cave, Nong Bua Lam Phu.