Coelosynapha loici Kjaerandsen, Polevoi & Salmela, 2020

Kjaerandsen, Jostein, Polevoi, Alexei & Salmela, Jukka, 2020, Coelosynapha, a new genus of the subfamily Gnoristinae (Diptera: Mycetophilidae) with a circumpolar, Holarctic distribution, Biodiversity Data Journal 8, pp. 54834-54834 : 54834

Coelosynapha loici Kjaerandsen, Polevoi & Salmela, 2020 sp. n.

Materials

Type status: Holotype. Occurrence: catalogNumber: TSZD-JKJ-102389 ; recordedBy: J. Kjaerandsen; individualCount: 1; sex: male; lifeStage: imago; preparations: Pinned, HMDS-dried from ethanol; occurrenceID: urn:uuid:5d9ce586-8e3f-4c53-b57c-4ce8f09debc9; Location: locationID: N-TRI-0045; country: Norway; stateProvince: Troms og Finnmark; municipality: Nordreisa; locality: Nokinivat, Reisa NP btw Naustneset and Nedrefosshytta ; verbatimElevation: 63 m; locationRemarks: MT 6-2016, pine forest; verbatimCoordinates: 69 19 38 N 21 57 18 E; decimalLatitude: 69.3273; decimalLongitude: 21.9551; geodeticDatum: WGS84; coordinateUncertaintyInMeters: 10 m; georeferenceProtocol: GPS; Identification: identifiedBy: J. Kjaerandsen; dateIdentified: Oct-07-2016; Event: samplingProtocol: Malaise trap; eventDate: Jul 16-Sep 16 2016; Record Level: collectionID: TMU-JKJ-COL-000389; institutionCode: TSZ; collectionCode: TSZD; ownerInstitutionCode: Tromso University Museum; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: catalogNumber: TSZD-JKJ-102390 ; recordedBy: J. Kjaerandsen; individualCount: 1; sex: male; lifeStage: imago; preparations: Pinned, HMDS-dried from ethanol, terminalia detached, cleared and stored in glycerine vial; occurrenceID: urn:uuid:b5f8e15a-36bb-488d-8487-db9c007600d0; Location: locationID: N-TRI-0045; country: Norway; stateProvince: Troms og Finnmark; municipality: Nordreisa; locality: Nokinivat, Reisa NP btw Naustneset and Nedrefosshytta ; verbatimElevation: 63 m; locationRemarks: MT 6-2016, pine forest; verbatimCoordinates: 69 19 38 N 21 57 18 E; decimalLatitude: 69.3273; decimalLongitude: 21.9551; geodeticDatum: WGS84; coordinateUncertaintyInMeters: 10 m; georeferenceProtocol: GPS; Identification: identifiedBy: J. Kjaerandsen; dateIdentified: Oct-07-2016; Event: samplingProtocol: Malaise trap; eventDate: Jul 16-Sep 16 2016; Record Level: collectionID: TMU-JKJ-COL-000389; institutionCode: TSZ; collectionCode: TSZD; ownerInstitutionCode: Tromso University Museum; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: catalogNumber: TSZD-JKJ-102391 ; recordedBy: J. Kjaerandsen; individualCount: 1; sex: male; lifeStage: imago; preparations: Pinned, HMDS-dried from ethanol, terminalia detached, cleared and stored in glycerine vial; occurrenceID: urn:uuid:69b453cf-4f75-41e2-8667-41c5fa37eadd; Location: locationID: N-TRI-0045; country: Norway; stateProvince: Troms og Finnmark; municipality: Nordreisa; locality: Nokinivat, Reisa NP btw Naustneset and Nedrefosshytta ; verbatimElevation: 63 m; locationRemarks: MT 6-2016, pine forest; verbatimCoordinates: 69 19 38 N 21 57 18 E; decimalLatitude: 69.3273; decimalLongitude: 21.9551; geodeticDatum: WGS84; coordinateUncertaintyInMeters: 10 m; georeferenceProtocol: GPS; Identification: identifiedBy: J. Kjaerandsen; dateIdentified: Oct-07-2016; Event: samplingProtocol: Malaise trap; eventDate: Jul 16-Sep 16 2016; Record Level: collectionID: TMU-JKJ-COL-000389; institutionCode: TSZ; collectionCode: TSZD; ownerInstitutionCode: Tromso University Museum; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: catalogNumber: TSZD-JKJ-102749 ; recordedBy: J. Kjaerandsen; individualCount: 1; sex: male; lifeStage: imago; preparations: Pinned, HMDS-dried from ethanol; occurrenceID: urn:uuid:4fe1af9c-e93a-4da7-8a17-1164022e4302; Location: locationID: N-TRI-0045; country: Norway; stateProvince: Troms og Finnmark; municipality: Nordreisa; locality: Nokinivat, Reisa NP btw Naustneset and Nedrefosshytta ; verbatimElevation: 63 m; locationRemarks: MT 6-2016, pine forest; verbatimCoordinates: 69 19 38 N 21 57 18 E; decimalLatitude: 69.3273; decimalLongitude: 21.9551; geodeticDatum: WGS84; coordinateUncertaintyInMeters: 10 m; georeferenceProtocol: GPS; Identification: identifiedBy: J. Kjaerandsen; dateIdentified: Jan-06-2017; Event: samplingProtocol: Malaise trap; eventDate: Jul 16-Sep 16 2016; Record Level: collectionID: TMU-JKJ-COL-000389; institutionCode: TSZ; collectionCode: TSZD; ownerInstitutionCode: Tromso University Museum; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: catalogNumber: TSZD-JKJ-102750 ; recordedBy: J. Kjaerandsen; individualCount: 1; sex: male; lifeStage: imago; preparations: Pinned, HMDS-dried from ethanol; occurrenceID: urn:uuid:a01d62c3-4453-4d80-9924-1e66531a6e41; Location: locationID: N-TRI-0045; country: Norway; stateProvince: Troms og Finnmark; municipality: Nordreisa; locality: Nokinivat, Reisa NP btw Naustneset and Nedrefosshytta ; verbatimElevation: 63 m; locationRemarks: MT 6-2016, pine forest; verbatimCoordinates: 69 19 38 N 21 57 18 E; decimalLatitude: 69.3273; decimalLongitude: 21.9551; geodeticDatum: WGS84; coordinateUncertaintyInMeters: 10 m; georeferenceProtocol: GPS; Identification: identifiedBy: J. Kjaerandsen; dateIdentified: Jan-06-2017; Event: samplingProtocol: Malaise trap; eventDate: Jul 16-Sep 16 2016; Record Level: collectionID: TMU-JKJ-COL-000389; institutionCode: TSZ; collectionCode: TSZD; ownerInstitutionCode: Tromso University Museum; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: catalogNumber: TSZD-JKJ-102751 ; recordedBy: J. Kjaerandsen; individualCount: 1; sex: male; lifeStage: imago; preparations: Pinned, HMDS-dried from ethanol; occurrenceID: urn:uuid:a01e5902-a3bd-4b97-8196-de178527d292; Location: locationID: N-TRI-0045; country: Norway; stateProvince: Troms og Finnmark; municipality: Nordreisa; locality: Nokinivat, Reisa NP btw Naustneset and Nedrefosshytta ; verbatimElevation: 63 m; locationRemarks: MT 6-2016, pine forest; verbatimCoordinates: 69 19 38 N 21 57 18 E; decimalLatitude: 69.3273; decimalLongitude: 21.9551; geodeticDatum: WGS84; coordinateUncertaintyInMeters: 10 m; georeferenceProtocol: GPS; Identification: identifiedBy: J. Kjaerandsen; dateIdentified: Jan-06-2017; Event: samplingProtocol: Malaise trap; eventDate: Jul 16-Sep 16 2016; Record Level: collectionID: TMU-JKJ-COL-000389; institutionCode: TSZ; collectionCode: TSZD; ownerInstitutionCode: Tromso University Museum; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: catalogNumber: TSZD-JKJ-102752 ; recordedBy: J. Kjaerandsen; individualCount: 1; sex: male; lifeStage: imago; preparations: Pinned, HMDS-dried from ethanol; occurrenceID: urn:uuid:49a80161-b4e1-4619-affb-88c05a2b6ae8; Location: locationID: N-TRI-0045; country: Norway; stateProvince: Troms og Finnmark; municipality: Nordreisa; locality: Nokinivat, Reisa NP btw Naustneset and Nedrefosshytta ; verbatimElevation: 63 m; locationRemarks: MT 6-2016, pine forest; verbatimCoordinates: 69 19 38 N 21 57 18 E; decimalLatitude: 69.3273; decimalLongitude: 21.9551; geodeticDatum: WGS84; coordinateUncertaintyInMeters: 10 m; georeferenceProtocol: GPS; Identification: identifiedBy: J. Kjaerandsen; dateIdentified: Jan-06-2017; Event: samplingProtocol: Malaise trap; eventDate: Jul 16-Sep 16 2016; Record Level: collectionID: TMU-JKJ-COL-000389; institutionCode: TSZ; collectionCode: TSZD; ownerInstitutionCode: Tromso University Museum; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: catalogNumber: TSZD-JKJ-102753 ; recordedBy: J. Kjaerandsen; individualCount: 1; sex: male; lifeStage: imago; preparations: Pinned, HMDS-dried from ethanol; occurrenceID: urn:uuid:6d2863e8-c396-4188-856d-fc3ea8d4575d; Location: locationID: N-TRI-0045; country: Norway; stateProvince: Troms og Finnmark; municipality: Nordreisa; locality: Nokinivat, Reisa NP btw Naustneset and Nedrefosshytta ; verbatimElevation: 63 m; locationRemarks: MT 6-2016, pine forest; verbatimCoordinates: 69 19 38 N 21 57 18 E; decimalLatitude: 69.3273; decimalLongitude: 21.9551; geodeticDatum: WGS84; coordinateUncertaintyInMeters: 10 m; georeferenceProtocol: GPS; Identification: identifiedBy: J. Kjaerandsen; dateIdentified: Jan-06-2017; Event: samplingProtocol: Malaise trap; eventDate: Jul 16-Sep 16 2016; Record Level: collectionID: TMU-JKJ-COL-000389; institutionCode: TSZ; collectionCode: TSZD; ownerInstitutionCode: Tromso University Museum; basisOfRecord: PreservedSpecimen Type status: Other material. Occurrence: catalogNumber: TSZD-JKJ-111229 ; recordedBy: J. Kjaerandsen; individualCount: 1; sex: male; lifeStage: imago; preparations: In 80% alcohol, in freezer; occurrenceID: urn:uuid:00150511-d1f9-4d15-8c76-cf7c00a1c58d; Location: locationID: N-TRI-0045; country: Norway; stateProvince: Troms og Finnmark; municipality: Nordreisa; locality: Nokinivat, Reisa NP btw Naustneset and Nedrefosshytta ; verbatimElevation: 63 m; locationRemarks: MT 6-2016, pine forest; verbatimCoordinates: 69 19 38 N 21 57 18 E; decimalLatitude: 69.3273; decimalLongitude: 21.9551; geodeticDatum: WGS84; coordinateUncertaintyInMeters: 10 m; georeferenceProtocol: GPS; Identification: identifiedBy: J. Kjaerandsen; dateIdentified: Feb-28-2020; Event: samplingProtocol: Malaise trap; eventDate: Jul 16-Sep 16 2016; Record Level: collectionID: TMU-JKJ-COL-000389; institutionCode: TSZ; collectionCode: TSZD; ownerInstitutionCode: Tromso University Museum; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: catalogNumber: MYCE-NV-2013-0230 ; recordedBy: J. Salmela; individualCount: 1; sex: male; lifeStage: imago; preparations: In 80% alcohol, terminalia detached, cleared and stored in glycerine vial; occurrenceID: urn:uuid:b0b3b9a8-f241-4219-848a-12bd65866d97; Location: country: Finland; stateProvince: Lapponia kemensis pars occidentalis; municipality: Kittilä; locality: Pomokaira, Tarpomapaeae ; verbatimElevation: 300 m; locationRemarks: Old-growth spruce forest; decimalLatitude: 67.8205; decimalLongitude: 25.9192; geodeticDatum: WGS84; coordinateUncertaintyInMeters: 10 m; georeferenceProtocol: GPS; Identification: identifiedBy: J. Salmela; Event: samplingProtocol: Malaise trap; eventDate: June 1-29 2009; Record Level: collectionID: MYCE-NV-2013-0230; institutionCode: LMM; collectionCode: LMM; ownerInstitutionCode: Regional Museum of Lapland; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: recordedBy: J. Salmela; individualCount: 1; sex: male; lifeStage: imago; preparations: In 80% alcohol; occurrenceID: urn:uuid:98e5f84a-192c-4750-bf98-ddbe1006bf4f; Location: country: Finland; stateProvince: Lapponia kemensis pars occidentalis; municipality: Kittilä; locality: Pomokaira, Tarpomapaeae ; verbatimElevation: 300 m; locationRemarks: Old-growth spruce forest; decimalLatitude: 67.8205; decimalLongitude: 25.9192; geodeticDatum: WGS84; coordinateUncertaintyInMeters: 10 m; georeferenceProtocol: GPS; Identification: identifiedBy: J. Salmela; Event: samplingProtocol: Malaise trap; eventDate: June 1-29 2009; Record Level: institutionCode: CSCA; collectionCode: CSCA; ownerInstitutionCode: California State Collection of Arthropods; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: catalogNumber: DIPT-JS-2015-0033 ; recordedBy: J. Salmela; individualCount: 1; sex: male; lifeStage: imago; preparations: In 80% alcohol; occurrenceID: urn:uuid:3fb6eaa2-3d60-4750-93e7-d0dbbdf55b49; Location: country: Finland; stateProvince: Lapponia enontekioensis; municipality: Enontekiö; locality: Pallas-Yllaestunturi National Park, Roeyninkuru ; verbatimElevation: 360 m; locationRemarks: Old-growth spruce forest, herb-rich; decimalLatitude: 68.1482; decimalLongitude: 24.075; geodeticDatum: WGS84; coordinateUncertaintyInMeters: 10 m; georeferenceProtocol: GPS; Identification: identifiedBy: J. Salmela; Event: samplingProtocol: Malaise trap; eventDate: Jul 5-Aug 7 2013; Record Level: collectionID: DIPT-JS-2015-0033; institutionCode: LMM; collectionCode: LMM; ownerInstitutionCode: Regional Museum of Lapland; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: catalogNumber: DIPT-JS-2014-0030 ; recordedBy: J. Salmela; individualCount: 1; sex: male; lifeStage: imago; preparations: In 80% alcohol; occurrenceID: urn:uuid:cd2dc33f-5eeb-4318-8067-e3b856374ff8; Location: country: Finland; stateProvince: Regio kuusamoensis; municipality: Salla; locality: Vaerrioe Strict Nature Reserve, Kuntasjoki ; verbatimElevation: 330 m; locationRemarks: Old-growth spruce forest, herb-rich; decimalLatitude: 67.7494; decimalLongitude: 29.6169; geodeticDatum: WGS84; coordinateUncertaintyInMeters: 10 m; georeferenceProtocol: GPS; Identification: identifiedBy: J. Salmela; Event: samplingProtocol: Malaise trap; eventDate: Jun 29-Jul 29 2013; Record Level: collectionID: DIPT-JS-2014-0030; institutionCode: LMM; collectionCode: LMM; ownerInstitutionCode: Regional Museum of Lapland; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: catalogNumber: DIPT-JS-2015-0151 ; recordedBy: J. Salmela; individualCount: 1; sex: male; lifeStage: imago; preparations: In 80% alcohol; occurrenceID: urn:uuid:25fc9a52-bb58-4f8c-8244-1f6437742075; Location: country: Finland; stateProvince: Lapponia enontekioensis; municipality: Enontekiö; locality: Pallas-Yllaestunturi National Park, Roeyninkuru ; verbatimElevation: 360 m; locationRemarks: Old-growth spruce forest, herb-rich; decimalLatitude: 68.1482; decimalLongitude: 24.075; geodeticDatum: WGS84; coordinateUncertaintyInMeters: 10 m; georeferenceProtocol: GPS; Identification: identifiedBy: J. Salmela; Event: samplingProtocol: Malaise trap; eventDate: Jul 5-Aug 7 2013; Record Level: collectionID: DIPT-JS-2015-0151; institutionCode: LMM; collectionCode: LMM; ownerInstitutionCode: Regional Museum of Lapland; basisOfRecord: PreservedSpecimen Type status: Other material. Occurrence: catalogNumber: DIPT-JS-2015-0393 ; recordedBy: J. Salmela; individualCount: 1; sex: male; lifeStage: imago; preparations: In 80% alcohol; Location: country: Finland; stateProvince: Lapponia kemensis pars orientalis; municipality: Savukoski; locality: Ainijaervi ; verbatimElevation: 260 m; locationRemarks: Old-growth spruce forest, herb-rich; decimalLatitude: 67.7701; decimalLongitude: 29.4332; geodeticDatum: WGS84; coordinateUncertaintyInMeters: 10 m; georeferenceProtocol: GPS; Identification: identifiedBy: J. Salmela; Event: samplingProtocol: Malaise trap; eventDate: Jul 30-Sep 28 2015; Record Level: collectionID: DIPT-JS-2015-0393; institutionCode: LMM; collectionCode: LMM; ownerInstitutionCode: Regional Museum of Lapland; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: catalogNumber: DIPT-JS-2016-0197 ; recordedBy: J. Salmela; individualCount: 1; sex: male; lifeStage: imago; preparations: In 80% alcohol; Location: country: Finland; stateProvince: Lapponia kemensis pars orientalis; municipality: Savukoski; locality: Ainijaervi ; verbatimElevation: 260 m; locationRemarks: Old-growth spruce forest; decimalLatitude: 67.7688; decimalLongitude: 29.4286; geodeticDatum: WGS84; coordinateUncertaintyInMeters: 10 m; georeferenceProtocol: GPS; Identification: identifiedBy: J. Salmela; Event: samplingProtocol: Malaise trap; eventDate: Jul 30-Sep 28 2015; Record Level: collectionID: DIPT-JS-2016-0197; institutionCode: LMM; collectionCode: LMM; ownerInstitutionCode: Regional Museum of Lapland; basisOfRecord: PreservedSpecimen Type status: Other material. Occurrence: catalogNumber: DIPT-JS-2016-0599 ; recordedBy: J. Salmela; individualCount: 1; sex: male; lifeStage: imago; preparations: In 80% alcohol; Location: country: Finland; stateProvince: Lapponia kemensis pars orientalis; municipality: Savukoski; locality: Ainijaervi ; verbatimElevation: 260 m; locationRemarks: Old-growth spruce forest; decimalLatitude: 67.7688; decimalLongitude: 29.4286; geodeticDatum: WGS84; coordinateUncertaintyInMeters: 10 m; georeferenceProtocol: GPS; Identification: identifiedBy: J. Salmela; Event: samplingProtocol: Malaise trap; eventDate: Jun 30-Jul 25 2015; Record Level: collectionID: DIPT-JS-2016-0599; institutionCode: LMM; collectionCode: LMM; ownerInstitutionCode: Regional Museum of Lapland; basisOfRecord: PreservedSpecimen Type status: Other material. Occurrence: recordedBy: A. Polevoi; individualCount: 1; sex: male; lifeStage: imago; preparations: Pinned, terminalia detached, cleared and stored in glycerine vial; occurrenceID: urn:uuid:2db6d72a-a309-406d-8917-6e1ee5e2f15d; Location: country: Russia; stateProvince: Murmansk Province; locality: Laplandskiy Nature Reserve, Lisiy Ruchei ; verbatimElevation: 237 m; decimalLatitude: 67.6512; decimalLongitude: 32.5985; geodeticDatum: WGS84; coordinateUncertaintyInMeters: 10 m; georeferenceProtocol: GPS; Identification: identifiedBy: A. Polevoi; Event: samplingProtocol: Malaise trap; eventDate: May 28-Sep 20 2014; Record Level: institutionCode: FRIP; ownerInstitutionCode: Forest Research Institute KRC RAS; basisOfRecord: PreservedSpecimen Type status: Paratype. Occurrence: recordedBy: A. Barkalov; individualCount: 1; sex: male; lifeStage: imago; preparations: Pinned, terminalia detached, cleared and stored in glycerine vial; occurrenceID: urn:uuid:a388b08d-4177-443f-be4e-1aa92cfe4077; Location: country: Russia; stateProvince: Chukotka Autonomyus District; locality: Anadyr r., 30 km lower l. Krasnoe ; verbatimElevation: 15 m; decimalLatitude: 64.72; decimalLongitude: 175.21; geodeticDatum: WGS84; coordinateUncertaintyInMeters: 1 km; Identification: identifiedBy: A. Polevoi; Event: samplingProtocol: Yellow pan trap; eventDate: Jun 25-Jul 19 2014; Record Level: institutionCode: SZM; ownerInstitutionCode: Siberian Zoological Museum; basisOfRecord: PreservedSpecimen Type status: Other material. Occurrence: recordedBy: Ju. Timchuk; individualCount: 2; sex: male; lifeStage: imago; preparations: In 80% alcohol, terminalia detached, cleared and stored in glycerine vial; occurrenceID: urn:uuid:b8160746-46c3-4fdc-94ea-d98b0ce9064f; Location: country: Russia; stateProvince: Republic of Altai; locality: 7 km NNE of Kurai ; verbatimElevation: 2144 m; decimalLatitude: 50.2957; decimalLongitude: 87.902; geodeticDatum: WGS84; coordinateUncertaintyInMeters: 10 m; georeferenceProtocol: GPS; Identification: identifiedBy: A. Polevoi; Event: samplingProtocol: Window trap; eventDate: May 28-Sep 20 2014; Record Level: institutionCode: UTR; ownerInstitutionCode: Tomsk State University; basisOfRecord: PreservedSpecimen

Description

Male. (Figs 3, 4, 6, 9, 10, 11 a, b, 12, n = 8)

Head (Fig. 4) dark brown, vertex sparsely covered by short, light hairs. Three ocelli almost in line, lateral ocellus placed 2.5 to 3 times its diameter from eye margin. Clypeus bud-shaped, 2x wider than long. Palpi 5-segmented, yellowish-brown. First three segments short, fifth segment 1.75-1.5 as long as fourth segment. Antenna with scape and pedicel usually largely brown, sometimes yellowish. Pedicel with outstanding apico-dorsal seta about as long as first flagellomere. Flagellomeres bearing light, curved, decumbent setulae, their length not exceeding width of respective flagellomeres.

Thorax (Fig. 4 a, c). Scutum usually dark-brown (with yellow humeral area in Chukotka specimen), thinly dusted, with yellow setulae. Pleural sclerites glabrous, dark brown, antepronotum and proepisternum yellowish. Laterotergite and mediotergite bare. Strong antepronotal seta about as long as head. Scutellum with two strong setae and four small setae.

Wings (Fig. 6). Wing length 2.6-2.9 mm. Sc ending in C, cross-vein sc-r absent. C produced beyond apex of R5 for about 2/3 of the distance between R4+5 and M1, R2+3 present, forming rectangular cell, occasionally separate veins R2 and R3 developed. Anterior fork long, divergent. Posterior fork short, its long stem is forked well beyond point of furcation of anterior fork. Veins M4 and CuA rather widely divergent. Sc with 3-5 macrotrichia on apical half. Basal transversal crossvein (tb) and M -stem bare, all other veins anterior of iCu fold setose. Halters pale yellowish.

Legs. Legs yellow with a brown tinge on trochanter, tibia and tarsi. Fore tarsus 1 slightly longer than fore tibia (ratio 1.06 in holotype).

Abdomen. Abdominal tergites and sternites brown, bearing light hairs. Tergites 7 and 8 rudimentary in size, sternite 8 lingulate.

Terminalia (Figs 9, 10, 11 a, b, 12). Yellowish-brown with pale yellow gonostyles except on sclerotised, blackish apical processes, setation mostly pale. Tergite 9 elongated, narrowed to half width on apical half, apex rounded, bearing 12-15 apico-ventral, stout setae. Cerci and hypoproct/sternite 10 partly retracted in under tergite 9, expandable. Gonocoxites with a deep ventral cleft. Gonocoxite with a hyaline, apico-mesal, long seta, almost as long as gonostylus. Gonostylus elongated, with three blunt, digitate projections apically (marked as a, b & c in Fig. 12 a, c), three lamellae subapically (marked as d, e & f in Fig. 12 a, b, c) and two long bristles mesally (marked as g & h in Fig. 12 b, c). Apical projection (a) finger-like, apically rounded, widest medially and bent dorsally. Subapical projection (b) wide, black, apically rounded, glabrous. Basal projection (c) black, hollow, having reticulate surface pattern, glabrous, apically rounded. Lateral lamella (d) triangular, high and narrow, glabrous. Ventral lamella (e) a sharp low ridge, edged with 6 blunt teeth-shaped setae. Ventromesal lamella (f) forming a blunt ridge densely covered with setulae.

Aedeagus inconspicuous, parameres fused.

Female unknown.

Etymology

Coelosynapha loici sp. n. is named in honour of the late French entomologist Loïc Matile (1938-2000) who first studied the new species and illustrated its terminalia (Fig. 1 ) way back in the mid 1980s. It is further an expression of the importance of classical morphological studies including detailed illustrations to be retained in the new integrative science of taxonomy.

Distribution

The new species has a wide Palaearctic range in boreo-mountainous localities, it has been collected from the High North boreal forest of Fennoscandia (Norway, Finland, NW Russia), from a high mountain site in Asian part of Russia and from hummock tundra in Far East Russia, at a total of seven localities (Fig. 2 ).

Ecology

Six of the collecting sites are within the boreal vegetation zone while the record from Chukotka is from the low arctic ecoregion. The European sites, one in Norway, three in Finland, one in Russia, are all northern boreal, close to the range limit of spruce ( Picea abies ssp. obovata ), lying some 140-180 km north of the arctic circle. All European collecting sites are pristine, boreal old-growth forests, dominated by spruce or occasionally pine (Reisadalen NP) and with scattered deciduous trees (birch, Betula pubescens and goat willow, Salix caprea ). The ground layer is characterised by mosses, especially Pleurozium schreberi and bilberry ( Vaccinium myrtillus ). Malaise traps on these sites have been set in the vicinity of springs or cold headwater streams. The specimens from Altai were collected at an altitude of 2144 m above sea level in steppe type Larix forest. The Chukotka specimen was collected with a yellow pan trap set in hummock tundra along the Anadyr river at 5-10 m above sea level.

Figures

Figure 1. Some 25 year ago, the late French entomologist Loic Matile (1938 - 2000) prepared these illustrations and a brief one-page description of a potential new genus based on a single specimen collected in Finland (A = ventral view, B = dorsal view.)

Figure 2. The circumpolar distribution of Coelosynapha gen. n. visualised on Google Earth. Green plots across the Palaearctic region display records of Coelosynapha loici sp. n., yellow plots across the Nearctic region display records of Coelosynapha heberti sp. n.