Thysananthus gottschei var. gottschei

Sukkharak, Phiangphak, 2015, A systematic monograph of the genus Thysananthus (Lejeuneaceae, Marchantiophyta), Phytotaxa 193 (1), pp. 448-450 : 448-450

8a. Thysananthus gottschei var. gottschei

Keel without appendage. Lobule free margin continuing into the ventral lobe margin but not into the margin of the dorsal lobe. Underleaf apex plane or recurved. Fig. 17 .

Distribution and ecology:— Indomalesia; 20–2100 m; on road-side trees, bark of living trees and fallen trees in lowland rain forests (primary and secondary forests) and montane forests. Fig. 5M .

Representative specimens:—India. ANDAMAN AND NICOBAR ISLANDS: Andaman islands, Port Blair, July 1890, Man s.n., ex hb. Levier 79 (G, PC) . Thailand. NAKHON SI THAMMARAT: San Yen, Sukkharak 758, 810 ( BKF, GOET) . Malaysia. PAHANG: on Genting Highland road, Sukkharak 717, 718, 720 ( BKF, GOET) ; Robinson hill, Ibrahim 36 ( KLU) .— PENANG: Penang hill, 11 February 1949, Allen s.n. ( BM) .— SABAH: Kinabatangan, Bukit Garam, Wood 1432 ( BM 2 packets, JE); Tawau , Bombay Burmeh Trading Co. Concession area , Wood 1389 ( BM) ;

32 • Phytotaxa 193 (1) © 2015 Magnolia Press

SUKKHARAK

Mt. Kinabalu , summit trail, Sukkharak 616 ( BCU) .— WESTERN NEW GUINEA: West Papua , Soron, 1872, Beccari s.n. ( JE) . Indonesia. JAVA: West Java , Tjibodas , 27 June 1898, Nyman s.n. (C 2 packets, G, S, W).— SULAWESI: Central Sulawesi , Lake Kalimpaa , Lore Lindu National Park, Gradstein 10324/2 ( GOET) .— SUMATRA: Bengkulu , Enggano Islands, “Modigliani”139 (G 2 packets, PC) . Philippines. NEGROS ORIENTAL: Cuernos Mts., May 1908, collector unknown. ( BM, G 3 packets, JE, W).— QUEZON: Polillo, McGregor 2–39, Bureau of science no. 10494 ( BM 2 packets). Papua New Guinea. CENTRAL: Kuriva logging area, Streimann & Vinas 14391 ( LAE) ; near Dabamura on Ower’s Corner road, Streimann & Naoni 14900, 14904, 14961 ( JE, LAE) ; road to Musgrave river , 9°36' S 147°38' E, Streimann & Naoni 15189 ( JE) ; Musgrave river road, Itikinumu, Streimann & Naoni 16690 ( JE, LAE) .— EASTERN HIGHLANDS: 25 miles S of Kainantu, Kurokawa 4370, 4371 ( JE) .— GULF: Bema-Kaintiba road, Streimann 33645 ( JE, LAE) .— MOROBE: logging area, 7°18' S 146°45' E, Streimann & Bellamy 13129 ( JE, LAE) ; 3 miles south-east of Bulolo , Shea 9097 ( NICH) ; Manki Trig. , Bulolo-Watut divide, Streimann & Bellamy 12997 ( JE, LAE) ; Slate-Gumi creeks divide, Streimann 13832 ( JE, LAE) ; upper Watut river, Streimann 23071, 23101 ( LAE) ; Herzog Mts. , Streimann & Umba 10983 ( LAE) ; Ekuti divide, Bulolo-Aseki road, Streimann 22623 ( JE, LAE) , Streimann 22629 ( NICH) ; Gumi divide, head of Gumi creek , Streimann 25156 ( JE, W); Sattelberg , June 1899, Nyman s.n. (C).— WEST NEW BRITAIN: South Bunga T. R. P., Bellamy 1357 ( JE, LAE, S); Evili river, Streimann 41598, 41608, 41609 ( JE) ; Geleo-Lasilai logging area at Mt. Laliti, Streimann 41088, 41209, 41253/a ( JE) , 41237 (JE, LAE, NICH), 41055, 41176, 41217 (JE, NICH).— WEST SEPIK: Aitape subdistrict, Darbyshire & Hoogland 8000 ( JE) ; Blackwater refugee camp, Mundua 314 ( JE, LAE) .

Reported from Sri Lanka by Eggers & Schäfer-Verwimp (1987) as Thysananthus convolutus .

Taxonomic notes:— Thysananthus gottschei was reduced to a synonym of T. convolutus and has been called the edentate form of the latter ( Grolle & Piippo 1984). Phytochemical and molecular results showed that this species should be re-instated ( Sukkharak et al. 2011 a, 2011b). The differences between T. gottschei and T. convolutus are discussed under the latter species.

Figures

FIGURE 5. Distribution of Thysananthus species. A. Thysananthus amazonicus (Spruce) Schiffn. B. Thysananthus spathulistipus (Reinwardt, Blume & Nees) Lindenb. C. Thysananthus convolutus Lindenb. D. Thysananthus mollis Steph., E. Thysananthus pancheri (Steph.) Hürl. F. Thysananthus anguiformis (Hook.f. & Taylor) Taylor ex Gottsche, Lindenb. & Nees G. Thysananthus comosus Lindenb. H. Thysananthus fruticosus (Lindenb. & Gottsche) Schiffn. I. Thysananthus aculeatus Herzog. J. Thysananthus appendiculatus Steph. K. Thysananthus montanus Gradst., X.-L. He & Piippo. L. Thysananthus retusus (Reinw., Blume & Nees) B.M.Thiers & Gradst. M. Thysananthus gottschei (Jack & Steph.) Steph. N. Thysananthus ciliaris comb. nov. O. Thysananthus discretus Sukkharak & Gradst.

FIGURE 17. Thysananthus gottschei (J.B.Jack & Steph.) Steph. A. Portion of shoot, ventral view. B. Leaves, ventral view. C. Underleaves, ventral view. D. Female bracts, ventral view: a = appendage. E. Female bracteole, ventral view. F. Perianth, dorsal view. G. Perianth, ventral view. H. Leaf apex. I. Margin cells of leaf lobe. J. Median cells of leaf lobe. K. Basal cells of leaf lobe. L. Leaf lobule apex. M. Cross section of stem. (A–C, H–M from the lectotype; D–G from Gradstein 10324 (GOET)).