Capoeta capoeta (Güldenstaedt, 1773)

Jouladeh-Roudbar, Arash, Ghanavi, Hamid Reza & Doadrio, Ignacio, 2020, Fig. 4 in Fig. 4 in Fig. 3 in Fig. 21. Sesarmops mora n in Paralbunea dayriti, Zoological Studies 59 (21), pp. 1-303 : 40-42

Capoeta capoeta (Güldenstaedt, 1773) – Native

( Fig. 66)

Cyprinus capoeta Güldenstädt [J. A. von] 1773: 508; Type locality: Tiflis, Caspian Sea. No types known.

Cyprinus fundulus Güldenstädt [J. A. von] 1787: 222; Type locality: Kura River at Tbilisi. No types known.

Scaphiodon asmussii Keyserling [E. von] 1861: 14 [17]; Type locality: Near Herat, Afghanistan. No types saved.

Capoeta sevangi De Filippi [F.] 1865: 312; Type locality: Lake Sevan (= Goktscha), Armenia. Holotype (unique): MZUT 695 .

Capoeta (Scaphiodon) steindachneri Kessler [K. F.] 1872: 49 [5]; Type locality: Zeravshan River, Uzbekistan. Syntypes: whereabouts unknown, not ZIN 4493–95 (1, 2, 3).

Capoeta hohenackeri Kessler [K. F.] 1877: 89; Type locality: Probably Kura-Araks river system, Azerbaijan. Holotype (unique): ZIN 2864 .

Capoeta gibbosa Nikolskii [A. M.] 1897: 344; Type locality: Bochsani [Bukhsani], se Khorasan, Iran. Syntypes: ZIN 11104 (2).

Common name: Pr: Siyah mahi ghafghazi, Siyah mahi mamoli, En: Caucasian scraper.

Diagnosis: This species is similar to C. aculeata and C. alborzensis but distinguished from them by a smaller scale size and a higher number of total lateral line scales (46–54 vs. 39–44).

Meristic characters: D: III–IV 7–8 (7), A: III 5, P: 16–20 (18–19), V: 9–10, GR: 19–21, LL: 55–58, TV: 43–44.

Distribution: Caspian Sea and Urmia Lake basins ( Fig. 67). Found in Aras, Balekhlochay, Qarasu, Kargan River in the Caspian Sea, also Gedar, Baranduz chay, Nazlo chay, Zarineh, Simineh and Ghale chay Rivers in the Urmia Lake basin.

Taxonomy: Berg (1949) placed in Varicorhinus but Coad (1991) transferred to Capoeta . Zareian et al. (2017) considered C. sevangi as a full and valid species without any clear justification; on the other hand, Ghanavi et al. (2016) using Cyt b genetic marker placed C. sevangi samples from Sevan Lake (Type locality of C. sevangi ) nested within C. capoeta samples from Arpa and Aras Rivers (see Fig. 1 in Levin et al. 2012 and Figs. 4–5 in Ghanavi et al. 2016). In the mentioned study, samples from the Sevan Lake, apart from not forming any independent clade, differed from other samples of C. capoeta with less than 0.4% genetic distance (closely related species in the same study had more than 1% genetic distance). In addition, we could not separate this species using the morphological diagnostic characters suggested by Zareian et al. (2017). Therefore, we consider C. sevangi as a synonym of C. capoeta , as suggested by previous authors ( Gabrielyan 2001; Turan et al. 2006; Ghanavi et al. 2016).

Conservation: IUCN: Least Concern ( Freyhof 2014e).

Figures

Fig. 1. Drainage basin map of Iran.

Fig. 4. Orders of freshwater fishes of Iran (Number of species, Percent of total).

Fig. 5. Families of freshwater fishes of Iran (Number of species, Percent of total).