Promalactis bilobignatha Wang, 2021

Wang, Shuxia & Liu, Chen, 2021, Taxonomic study of the genus Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) X. The commotica species-group, with descriptions of twenty-two new species, Zootaxa 4980 (2), pp. 293-330 : 301-302

Promalactis bilobignatha Wang , sp. nov.

( Figs 6 , 28 )

Type material. CHINA, Yunnan: Holotype ♂ , Lingbaoshan Forest Park, Nanjian , 2338 m, 25.VIII.2015, leg. KL Liu & H Wei, slide No. LC 19354 . Paratypes (2♂): same data as holotype, slide No. LC 19343 ♂ .

Diagnosis. The new species can be distinguished from its congeners by the large costal forewing spot from between distal 2/5 and 1/5 abruptly narrowed ventrad; and in the male genitalia by the bilobed uncus, the bilobed gnathos, and the symmetrical valva with a flagellate dorsoapical lobe that is 2.5 times the length of the ventroapical lobe.

Description. Adult ( Fig. 6 ). Forewing length 6.0‒ 7.5 mm.

Head: Vertex blackish brown, with snowy white scales laterally. Labial palpus with second palpomere orange yellow; third palpomere black except dirty white at apex. Antenna with scape dirty white on dorsal surface, pale grey on ventral surface; flagellum black annulated with white, except basal several flagellomeres dirty white on dorsal surface.

Thorax: Dorsum of mesothorax and tegula dark brown. Forewing yellowish brown to dark ochreous brown; markings white edged with blackish brown scales: costal spot large, inverted triangular, from between distal 2/5 and 1/5 narrowed ventrad and crossing anterior angle of cell; basal streak obliquely inward to base of fold; antemedian streak L-shaped, from dorsal 2/5 to basal 1/3 of anterior margin of cell, curved inward at anterior 2/5; dorsal streak from basal 3/5 obliquely outward to distal 1/5 of posterior margin of cell; tornal spot black, diffused inward to end of dorsal streak; pretornal spot represented by several white scales; apical patch consisting of three white spots, separated by black scales, narrowed from apex to posterior corner of termen; fringe orange yellow except greyish brown around tornus. Hindwing and fringe grey. Foreleg black, femur and tibia greyish white ventrally, tibia with a white spot at base and middle respectively, with a tuft of white scales at apex, tarsus white at apices of basal two tarsomeres; midleg black dorsally, yellowish brown ventrally, tibia mixed with white scales, with three white spots, with a tuft of white scales at apex, tarsus white at apices of basal two tarsomeres; hindleg yellowish brown ventrally, brown dorsally except distal four tarsomeres white.

Abdomen: Male genitalia ( Fig. 28 ). Uncus rectangular, sclerotized, and folded inward on basal half laterally; distal 1/3 concave semicircularly at middle on posterior margin, forming two stout lobes posterolaterally. Gnathos slightly longer than uncus; mesial plate squamous on distal 3/5, apex concave semicircularly at middle, forming two digitate apical lobes. Tegumen widened to posterior 1/3; lateral arms narrowed to obtuse apex. Valva almost parallel basally, bilobed distally; costa very short, produced to a long free flagelliform dorsoapical process nearly as long as valva, with fine spines along midline from base to apex, forming a longitudinal cluster, pointed at apex; sacculus wide at base, narrowed to middle, distal half extremely narrow to ventroapical process; ventroapical process wider, shorter than half length of dorsoapical process, arched on dorsal margin, roundly concave at middle on ventral margin, forming a right inner angle and an obtuse outer angle ventrally, apex rounded, with tiny denticles. Saccus triangular, as long as uncus, wide at base, narrowed to pointed apex. Juxta widened to basal 1/3, thereafter uniform to distal 1/3, then abruptly narrowed to pointed apex on right side, longitudinally with a tuft of fine spines extending from middle to apex; basal lobe clubbed, reaching before apex of saccus.Aedeagus shorter than valva, straight, with short spines and claw-like apically; cornutus absent.

Female. Unknown.

Distribution. China (Yunnan).

Etymology. The specific epithet is derived from the Latin bilobus and -gnathus, referring to the distally bilobed uncus and gnathos.

Figures

FIGURES 1‒8. Adults of Promalactis spp. 1, P. apiculifera, holotype, ♂; 2, P. apisetosa, holotype, ♂; 3, P. arcicosta, holotype, ♂; 4, P. baoshana, holotype, ♂; 5, P. bicornuta, holotype, ♂; 6, P. bilobignatha, paratype, ♂; 7, P. binipapillata, paratype, ♀; 8, P. bispina, paratype, ♀. Scales = 2 mm.

FIGURES 23‒28. Male genitalia of Promalactis spp. 23, P. apiculifera, holotype, slide No. HXM14207; 24, P. apisetosa, holotype, slide No. DZH11066; 25, P. arcicosta, paratype, slide No. LC19399; 26, P. baoshana, holotype, slide No. JYY17588; 27, P. bicornuta, holotype, slide No. DZH10246; 28, P. bilobignatha, paratype, slide No. LC19354, 28a, enlarged aedeagus. Scales = 0.2 mm.