Dardanus scutellatus (H. Milne Edwards, 1848 )

Malay, Maria Celia (Machel) D., Rahayu, Dwi Listyo & Chan, Tin-Yam, 2018, Hermit crabs of the genera Calcinus Dana, Clibanarius Dana, and Dardanus Paul’son from the PANGLAO 2004 Expedition, with description of a new species and a checklist of the hermit crabs of the Philippines (Crustacea: Anomura: Paguroidea), Raffles Bulletin of Zoology 66, pp. 23-65 : 51

Dardanus scutellatus (H. Milne Edwards, 1848)

( Fig. 4E )

Pagurus scutellatus H. Milne Edwards, 1848: 61 ( Zanzibar, type locality); Forest, 1953b: 560–561; Fize & Serène, 1955: 189–195, Fig. 29, pl. 6 nos. 5–10.

Pagurus fabimanus – Estampador, 1937: 503; Yap-Chiongco, 1938: 199; Estampador, 1959: 50.

Dardanus scutellatus – Gherardi & McLaughlin, 1994: 641–642.

Material examined. 1 female SL 8.8 mm ( ZRC), stn B7, Catarman, Panglao I., 9°35.9′N, 123°51.8′E, 4–30 m, 5.vi.2004 ; 1 male SL 6.0 mm ( ZRC), stn D5, Pontod Islet, Panglao I., 9°33.6′N, 123°43.5′E, 0–3 m, 6.vi.2004 ; 1 male SL 9.7 mm ( ZRC), stn M18, Gak-Ang Islet, off Panglao I., 9°33.0′N, 123°45.5′E, 0–1 m, 10 & 12.vi.2004 ; 1 male SL 9.8 mm ( ZRC), stn D12, Panglao Channel, Tagbilaran, Panglao I., 9°38.5′N, 123°51.0′E, 2–4 m, 20.vi.2004 ; 1 ovig. female SL 4.5 mm, 1 female SL 5.2 mm, 2 males SL 6.3–6.5 mm ( ZRC), stn M58, Balicasag I., 9°31.3′N, 123°41.0′E, depth unknown, 4.vii.2004 ; 2 ovig. females SL 5.1–7.7 mm, 5 males SL 4.7–8.9 mm ( ZRC), stn M51, Mayacabac, Panglao I., 9°36.8′N, 123°52.2′E, 0 m, 30.vi.2004 ; 5 females SL 3.1–9.1 mm, 5 males SL 4.0– 6.8 mm ( ZRC), stn R50, Looc, Panglao I., 9°35.7′N, 123°44.4′E, 3–7 m, 17.vi.2004 .

Habitat. Intertidal to shallow subtidal, 0 to approximately 5 m deep, seagrass beds and reef slopes, on sandy and soft substrates.

Remarks. This species resembles D. guttatus , but is distinctively different in colouration (see ‘remarks’ under latter species) and in habitat. It is also affiliated to D. woodmasoni , but can be distinguished by the lower margin of the big chela, which is smooth or poorly denticulated, while it is obviously denticulated in D. woodmasoni . Dardanus scutellatus is a relatively uncommon species.

Figures

Fig. 4. A, Dardanus sanguinolentus (Quoy & Gaimard, 1824). Female ovig. SL 6.3 mm, stn R6; B, Dardanus balhibuon, n. sp. Holotype female SL 14.3 mm, stn R38, NMCR 40109, preserved colouration; C, Dardanus megistos (Herbst, 1804). Male SL 11.0 mm, stn M8; D, Dardanus pedunculatus (Herbst, 1804). Male SL 13.4 mm, stn N3; E, Dardanus scutellatus. Male SL 9.8 mm, stn D12; F, Dardanus woodmasoni (Alcock, 1905). Male SL 5.8 mm, stn R76.