Tibiodrepanus sinicus ( Harold, 1868 )

Barbero, Enrico, Palestrini, Claudia & Roggero, Angela, 2011, Tibiodrepanus tagliaferrii — a new Afrotropical Drepanocerina species (Coleoptera: Scarabaeidae: Oniticellini), with notes on phylogeny and distribution of the genus, Zootaxa 2923, pp. 27-47 : 40-41

Tibiodrepanus sinicus ( Harold, 1868)

( Figs. 14 , 18, 22 , 26 )

Drepanocerus setosus sensu Boheman 1858 (nec Wiedemann 1823) Drepanocerus sinicus Harold, 1868

Type material. Holotype: 3, “Kinb. China ”, “ Type ”, “ setosus Bhm. ”, “ Typus (…)” ( NHRS).

Additional material examined. We examined over 200 specimens of T. sinicus , that are listed below, except for when the localities were considered too generic (see remarks for further information). The number of individuals for each locality is given within brackets.

BUTHAN: [1] Punaka, 5.x.2004, 1600 m, Bellini leg. ( SZCM). CHINA: [1] Canton, S. 1925 ( NHMB); [15] Hong Kong ( BMNH, OUMNH, ZMHB, ZSM); [2] same locality, 16.x.1849 ( OUMNH); [6] same locality, Dohrn leg. ( ZMHB, ZMUC); [1] same locality, Pascoe leg. ( BMNH); [9] same locality, Walker leg. ( BMNH); [4] Hong Kong, Northern Territories, 1996-98, De Rougemont leg. ( OUMNH); [2] Hupeh Province, Wu-schan Mts. ( ZMHB); [9] Fujian, Shaowu, 500 m, 30.x.1937, Klapperich leg. ( NMPC, HNHM, EBCT); [1] Fujian, Fuzhou [=Foo chow], 1862, [Lewis coll. 1915-88] ( BMNH); [1] Wallau ( ZMHB); [2] Yunnan, Yunxian [Yun Hsien], iii.1942, Jellison leg. ( USNM); [3] Yunnan, Jingdong county, 1200m, 4.iii.1957, Panfilon leg. ( ZIN); [1] same locality and collector, 5.iii.1957 ( ZIN); [18] Hainan, Sanya, 5.ii.1989, Majzlan leg. ( CSC); [3] SE China, Guangxi District, Yulin Liuwan Mts., SW Yulin, 200m, 18.xi.1993, Schillhammer leg. ( CSC); [1] Guangxi, 10 km N Liuzhou, 10.xi. 1993, 200m, Schillhammer leg. ( CSC); [1] Hainan, Wuzhishan, 18.i.1996, Jäch leg. ( CSC). INDIA: [1] Khandesh [near Mumbay], Bell leg. ( OUMNH); [1] Tamil Nadu [S. Indien], Kodaikanal, Pulney Hills, 6500 ft., iv.1953, Nathan leg. ( NHMB); [1] Goa, Molem, 20–22.ii.1980. Topál leg. ( HNHM); [1] Uttarakhand, Dehra Dun, Suawa River, xi.1930, Beeson leg. ( BMNH); [1] Uttarakhand, Tharali [=Uttar Prades India bor. Thalari], 24.vii.1994, Snizek leg. ( EBCT); [1] Uttarakhand [= India bor. Uttar Prades], Rishikesh, 340m 5.vii.1994, Snizek leg. ( EBCT); [1] Uttarakhand, Dhobi Ghat, Mussorie, Dehradun, 14.iv.1922, Bose leg. ( BMNH); [2] Himachal Pradesh [=W Himalaya], Manali env., 2200m, 1.vii.2003, Gorodimhi leg. ( ZIN). INDONESIA: [1] Java, Tjikidang [=Cikidang], 1902 ( CSH). LAOS: [1] Naphok, 27.iii.1990, Kondorosy leg. ( HNHM); [2] C Laos, Khammouan province, Ban Khoun Ngeun; 200m ca., 24–29.iv.2001, Pacholatko leg. ( NHMB); [1] CE Laos, Boli Kham Xai province, Ban Nape (8 km NE), 600m ca., 1–18.v.2001, Pacholatko leg. ( NHMB); [4] N Laos, Lg Nam Tha Province, ca. 30 km NW Lg Nam Tha, 800m, 18.vi.1996, Schillhammer leg. ( CSC). MALAYSIA: [1] Penang, 4– 9.i.1915, Cameron leg. ( BMNH); [9] same locality,. 6.i–1.ii.1981, Palm leg. ( MZLU); [2] same locality, 6.i– 1.ii.1981, Palm & Ferrinchi leg. ( MSNG). NEPAL: [1] Pokhara Sarangkot, 1592 m 15.xi.1998 ( CSH). PHILIP- PINES: [1] Luzon, Los Banos ( NHMB). SINGAPORE: [1] Singapore, Saunders ( BMNH); [1] Singapore, Scott Land, ii.1922, Saunders ( BMNH); [1] same locality and collector, iii.1922 ( BMNH); [1] same locality and collector, 31.xii.1922 ( BMNH); [1] same locality and collector, 5.xi.1922 ( BMNH). THAILAND: [3] Chiang Mai, Bantchetawan Lamphun, 5.vii.1990, Minetti leg. ( PMOC); [2] NM Thailand, Chom Thong, 24–27.iv.1991, Dembicky leg. ( NHMB); [1] same locality and date, Pacholatko leg. ( NHMB); [1] NW Thailand, Mae Hong Son, 7.v.1991, Dembicky leg.( NHMB); [25] Pattaya, 12.xii.1979, Palm leg. ( MZLU, MSNG); [1] Surin Province, Ta Moan temples [Ta-maon. I(I) 92, 87, 7386], border to Cambodia, 1.i.1992 ( BMNH). UNION OF MYANMAR: [1] Rangoon, v.1885, Fea ( MSNG); [14], Kayin Hills [= Carin], Asciuii Ghecù, 1400–1500m, iii–iv.1888, Fea leg. ( MSNG, MHNG, IRSNB, BMNH); [2] same locality and collector, 1200–1400m, iii.1888 ( BMNH); [1] Kayin Hills [= Carin], Chebà, 800–1000m, xii.1887, Fea ( MSNG). VIETNAM: [2], Tonkin, Hoa-Binh ( NHMB, MNRS); [3] same locality, 1928, de Cooman leg. ( IRSNB); [1], Luc Yen, 5.xii.1971, Topál leg. ( HNHM); [2], Yanyen (= Song Da), Rivière Noire, 1923-24, Dussault leg. ( NHMB); [17], Dong Nai Province, Vinh Cuu District, Tri An Lake, 60 m, 21.viii.2005, Mann leg. ( OUMNH); [2], N Vietnam, 52 km SW of Lang Son, 27.iv.–6.v. 1996, 370m, Pacholatko & Dembicky leg. ( NHMB); [7] S Vietnam, Nam Cat Tian National Park, 1–15.v.1994, Pacholatko & Dembicky ( CSC); [1] Quang Nam Da Nang and Gia Lai-Kon Tum Provinces border (near Laos border), 400– 1100m, 19–21.i.1997, Businsky & Businsky leg. ( ABCB); [1] Lam Dong Province, Baolac District, Baolac Pass, 800m, 20.viii.2005, Mann leg. ( OUMNH).

Diagnosis. The species is separated from the other Oriental species by the surface of the head densely, sparsely, very superficially punctate.

Distribution. Buthan, India, Nepal, China, Union of Myanmar, Laos, Thailand, Vietnam, Singapore, Philippines, Indonesia and Malaysia.

Literature. Drepanocerus setosus . Boheman 1858:50, Harold 1868:104. Drepanocerus sinicus . Harold 1868:104, Harold 1869:1041, Gillet 1911:100, Boucomont & Gillet 1921:20, Winkler 1924:1028, Boucomont 1923:3, Boucomont 1929:764, Arrow 1931:383, Paulian 1945:139, Janssens 1953:31, Balthasar 1963:68, Kryzhanovsky & Medvedev 1966:397, Masumoto 1976:1, Mittal 1981:37, Simonis 1985:295, Masumoto 1987:130, Majzlan & Rychlik 1991:520, Kabakov 1998:41, Kabakov & Napolov 1999:72, Chandra 2000:360, Bedzek & Krell 2006:156, Sewak 2009:36. Tibiodrepanus sinicus . Krikken 2009:21, Sabu et al. 2011:35.

Remarks. The species was erroneously identified as D. setosus by Boheman (1858), but later Harold (1868) identified it as D. sinicus . The species was included in Tibiodrepanus by Krikken (2009). Some localities could not be precisely georeferenced, thus they are not listed above [e.g., two specimens from northern China, Meyen (ZMHB), and one specimen from India (USNM)].

Figures

FIGURES 12 – 15. Epipharynx. Tibiodrepanus hircus: 12; T. setosus: 13; T. sinicus: 14; T. sulcicollis: 15. Scale bar = 0.2 mm.

FIGURES 16 – 23. Male genitalia. Tibiodrepanus hircus aedeagus: 16; T. setosus aedeagus: 17; T. sinicus aedeagus: 18; T. sulcicollis aedeagus: 19; Tibiodrepanus hircus endophallus laminae: 20; T. setosus endophallus laminae: 21; T. sinicus endophallus laminae: 22; T. sulcicollis endophallus laminae: 23. Aedeagus scale bar = 0.5 mm, endophallus laminae scale bar = 0.2 mm.

FIGURES 24 – 27. Female genitalia. Tibiodrepanus hircus vagina and receptaculum seminis: 24; T. setosus vagina and receptaculum seminis: 25; T. sinicus vagina and receptaculum seminis: 26; T. sulcicollis vagina and receptaculum seminis: 27. Scale bar = 0.2 mm.