Calvariopsis saopaulense, Ruta, 2019

Ruta, Rafał, 2019, Calvariopsis gen. nov., a new genus of Neotropical Scirtidae (Coleoptera: Scirtoidea), Zootaxa 4604 (1), pp. 1-41 : 32

Calvariopsis saopaulense sp. nov.

( Figs 1W , 25 )

Type material. Holotype, male ( NMPC): “ SAO PAULO \ BRAS MRÁZ LGT. \ MUS. PRAGENSE” . Paratype, male ( NMPC): same data as holotype .

Diagnosis. Small (TL 1.9–2.1 mm), blackish, widest in anterior portion of elytra, externally resembling C. wittmeri sp. nov. Identification is possible only on the basis of morphology of male genitalia: trigonium of penis very narrow, tegmen V-shaped (unique in known Calvariopsis gen. nov,), without basal apodemes, apical plate of tergite VIII with V-shaped emargination.

Description. Male. Body oblong oval, convex, covered with brownish, suberect setae. Body brown, legs, antennae and mouthparts yellowish. Head very broad, 1.7× wider than width of interocular space, with fine punctation, punctures small, granulate, separated by a 2.0–3.0× diameter of a puncture, eyes moderately large, not protuberant. Pronotum transverse, widest at its base. Disc of pronotum with fine, granulate punctation, punctures separated by ca. 2.0× diameter of a puncture. Posterior margin of pronotum subtly bisinuate. Anterior angles protruding anteriorly, subtriangular, posterior angles obtuse. Scutellar shield subtriangular, covered with subtle punctation. Elytra oblong, without traces of longitudinal ridges, widest slightly before the middle of their length, sides curved, regularly converging to apex in posterior 1/3. Humeri well marked. Punctation much stronger than on pronotum, punctures elongate, distance between punctures ca. the diameter of a puncture.

Male genitalia. Penis (L 0.45 mm, W 0.07 mm, Fig. 25A ) moderately wide, with slightly curved sides, param- eroids long, subtriangular, gradually narrowing, pointed at apices, trigonium very narrow, subtriangular. Tegmen (L 0.27 mm, W 0.16 mm, Fig. 25B ) V-shaped, apices of parameres pointed, covered with setae. Sternite IX (L 0.17 mm, W 0.11 mm, Fig. 25B ) small, subrectangular, with single, short apodeme, apical portion of sternite covered with numerous setae. Tergite VIII (L 0.25 mm, W 0.25 mm, Fig. 25C ) with V-shaped emargination of anterior margin, anterior margin with small denticles and small setae, apodemes as long as apical plate. Tergite IX (L 0.23 mm, W 0.21 mm, Fig. 25D ) with membranous and relatively narrow apical plate, apodemes as long as apical plate, strongly diverging.

Female. Unknown.

Measurements and ratios. Males (n = 2) TL 1.93–2.07 (2.00) mm, PL 0.40–0.43 (0.42) mm, PW 0.90–0.97 (0.93) mm, EL 1.73–1.83 (1.78) mm, EW 1.23–1.33 (1.28) mm, TL/EW 1.55–1.57 (1.56), PW/PL 2.23–2.25 (2.24), EL/EW 1.38–1.41 (1.39), EL/PL 4.23–4.33 (4.28).

Distribution. Known only from the type locality in Sao Paulo, Brazil.

Etymology. Named after Sao Paulo, where the type locality of the species is located.

Figures

FIGURE 1. Calvariopsis gen. nov., dorsal view. A) C. borowieci sp. nov., male; B) C. borowieci sp. nov., female; C) C. four- gassiense sp. nov., male; D) C. panamense sp. nov., male; E) C. peruviana sp. nov., Dept. Cusco, male; F) C. peruviana sp. nov., Callanga, male; G) C. peruviana sp. nov., Dept. Junín, male; H) C. picta sp. nov., Callanga, male; I) C. pittieri sp. nov., male; J) C. pittieri sp. nov., female; K) C. sculptipenne (Pic, 1931), female; L) C. venezuelense sp. nov., male; M–O) C. yanay- acuense sp. nov., males; P) C. yanayacuense sp. nov., female; Q) C. bituberculata sp. nov., male; R) C. brasiliense (Pic, 1916), male; S) C. brasiliense, female; T) C. guyanense sp. nov., male; U) C. kawense sp. nov., male; V) C. nana sp. nov., male; W) C. saopaulense sp. nov., male; X) C. wittmeri sp. nov., male; Y) C. sp., Bolivia, female. Scale bars = 1.0 mm.

FIGURE 25. Calvariopsis saopaulense sp. nov., male.A) penis; B) tegmen and sternite IX; C) tergite VIII; D) tergite IX. Scale bar = 0.25 mm.