Physarum bethelii T. Macbr. ex G. Lister, 1911

Lado, Carlos & Basanta, Diana Wrigley De, 2018, Typification of the myxomycete taxa described by the Listers and preserved at the Natural History Museum, London (BM), Phytotaxa 341 (1), pp. 448-450 : 448-450

Physarum bethelii T. Macbr. ex G. Lister , in Lister, Monogr. mycetozoa, ed. 2, 57. 1911

≡ Physarum viride var. bethelii (T. Macbr. ex G. Lister) Sturgis , Colorado Coll. Stud. Sci. Ser. 12(12): 439. 1913

— Lectotypus (designated here): U.S.A. Colorado, Colorado Springs, Cheyenne Mountain , 38º44’N 104º52’W, 10 Aug 1908, Dr. W.C. Sturgis WCS A 62 ( BM 2134 !, BM001089850 ) . Syntypus: U.S.A. Colorado, Colorado Springs, Cheyenne Mountain , 38º44’N 104º52’W, 10 Sep 1908, Dr. W.C. Sturgis WCS A 62 ( BM s.n.!, two boxes WCS A 62 BM001247131 , BM001247132 + one slide BM s.n.!, BM001223779 ) . Syntypus: U.S.A. Colorado, 1908?, E. Bethel ( BM 2133 !, Macb. 73, BM001089851 )

BM 2134 (WCS A 62)/ Physarum bethelii Macbr. / Cheyenne Mt./ Colorado Springs/ Colo./ Aug. 10.08 (St.2.128)/ BM001089850 [sheet label dated: 12 JUN. 1912: “BM 2134/ Physarum bethelii Macbr. in litt./ Cheyenne Mt., Colorado Springs/ 10.8.1908, leg. Dr. W.C. Sturgis”]. BM s.n. two boxes WCS A 62 / Physarum viride Pers. / var. bethelii Sturg. / Cheyenne Mt./ Colorado Springs/ Colo./ Sept.10.08 St. 2 p. 119/ BM001247131, BM001247132 + one slide, BM001223779. BM 2133 / Physarum bethelii Macbr. in litt./ Colorado, leg. E. Bethel, 1908?/ (Macb. 73)/ (NXXVIII.123)/ Botanical Laboratory State University of Iowa / Tilmadoche / bethellii/ n.s./ Colorado 08./ BM001089851. [Two revision labels, one by D.T. Kowalski July 1971 “ Type specimen”, and another by G. Adamonyte “Examined 21–11–2002 macroscop., no slides made.”].

NOTES: Lister (1911: 57) reported this species as “ P. bethelii Macbr. in litt.” and based her description on the American material “On dead wood.– Colorado ( BM 2133 , 2134 ).”. The author added “It has been found twice, each time in Colorado, once by Mr. E. Bethel, once by Dr. Sturgis ”. In BM both collections were located ( BM 2133 and BM 2134 ) and we designate the second as the lectotype, as it is in better condition and has notes and drawings in the notebook St.2: 128 where G.Lister kept of all specimens received from Sturgis. In notebook XXVIII: 123–124 there are notes on collection “Macb. 73” ( BM 2133 ). There were also 4 more boxes from the same place and two of them are labelled WCS [Sturgis] A62, and a slide with a reference to the Sturgis notebook (St.2: 119). These could all be duplicates of BM 2134 , that bears the number WCS A62 , but they are from a later date. The other boxes are labelled WCS 781 from Aug. 27.08 and Aug. 8.12, and have a reference to the Sturgis notebook (St. 2: 166, 167, 168) where we found notes and illustrations of this material .