Ayacucho tapacocha, Benedetti & Pinto-da-Rocha, 2022

Benedetti, Alipio Rezende & Pinto-da-Rocha, Ricardo, 2022, Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species, Arthropod Systematics & amp; Phylogeny 80, pp. 309-388 : 309

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Ayacucho tapacocha

nom. nov.

3.22. Ayacucho tapacocha nom. nov.

Figs 3C View Figure 3 , 7S, T View Figure 7 , 16 G-I View Figure 16 , 23C, D View Figure 23 , 28 View Figure 28

Tapacochana insignita Roewer, 1957: 73 (desc.), figs. 25 (dorsal habitus), 26 (pedipalpus); Kury 2003: 145 (cat.); Coronato-Ribeiro and Pinto-da-Rocha 2017: 203 (cit), 235-236 (mat).


MALE: Measurements (n =15) DSW: 3.3-5.1 (3.3); DSL: 4.0-5.6 (4.0); CL: 1.3-2.2 (1.3). FIVL: 6,2-6,5. ChL: 1.3-4.0 (1.3). Coloration: (Fig. 23C View Figure 23 ) Chelicerae, pedipalpus, carapace, lateral margins of DS reddish-brown. Areas I-IV, coxa IV, femora-tibiae I-IV dark brown (some specimens may have a longitudinal lighter strip running through the carapace and areas of dorsal scutum). Posterior margin of DS and free tergites I-III black. Dorsum: (Fig. 3C View Figure 3 ) Alpha-type DSS, with constriction II shallow (almost faint). Anterior margin of dorsal scutum with median elevation; granules sparsely distributed. Ocularium with a pair of spines. Carapace with granules scattered in the lateral and posterior regions. Areas I-IV densely granulate; one pair of median tubercles in all areas. Posterior margin of DS and free tergites I-III with a row of acuminate tubercles, higher than those on scutal areas. Lateral margins of DS with granules distributed throughout their length. Chelicerae: (Fig. 3C View Figure 3 ) In smaller males, chelicerae slightly larger than the females (as in the holotype). Swollen in large males. Segment I granulate in retrolateral half. Segment II predominantly smooth with one tooth on the finger. Segment III with two teeth. Pedipalpus : Small granules distributed on the dorsal surface of the trochanter, femur, patella and tibia. Trochanter with a ventrodistal setiferous tubercle. Femur with a ventrobasal setiferous tubercle; wit a row of five or six ventral setiferous tubercles, except in the apical portion. Tibia: prolateral IiII retrolateral iiIi. Tarsus: prolateral iIii, retrolateral iiiIiii. Venter: Coxa I with a median row of five small tubercles. Coxae II-IV with few granules. Genital area with few scattered granules. Free sternites with a row of small granules. Operculum anal with granules sparsely distributed on its surface. Legs: (Figs 3C View Figure 3 , 7S, T View Figure 7 ) Coxae I-III each one with two apophyses (one prolateral, one retrolateral). Coxa IV with granules distributed throughout its surface; a distal prolateral apophysis apically acuminated. Trochanters I-III smooth. Trochanter IV with three small dorsoapical tubercles and a small median retroapical tubercle, apically acuminated. Femora I-III with granules scattered throughout their length. Femur IV densely granulate; a retroventral row of 7-9 small acuminate tubercles along the distal ½, apically growing; a proventral row of tiny tubercles, growing apically, the last three being larger and apically curved; a retrolateral row of 11-14 tubercles along the distal ⅔; a prolateral row of 11-12 tubercles along the distal ⅔. The shape of the femur IV tubercles are extremely variable and may be with blunt apex, acuminate and straight; in all rows of tubercles, the basal ones are tiny and round, slightly larger than granules, whereas the others have varied morphology (apex blunt, acuminate or lanceolate). Patellae I-III with sparse granules. Patella IV dorsally densely granulate, with granules more sparsely distributed on ventral surface; with three smaller dorsoapical tubercles and one larger tubercle. Tibiae I-III with granules throughout their length; tibia IV densely granulate, with a row of 13-14 ventral acuminate tubercles, growing apically. Tarsal segmentation: (n =15) 7, 10-12 (10), 6-9 (6), 7-10 (7). Penis: (Fig. 16G-I View Figure 16 ) VP subrectangular, with lateral margins slightly convex (in ventral view); distal margin slightly convex; with laterodistal projections; MS C1-C4 subapical, long and curved; MS A1-A2 median long and straight; MS B1 sub basal very short; MS D1 median short and straight. Lateral sacs long, robust and with blunt apex. Stylus slightly flattened laterally, apically inflated; with a long dorsal projection, at a 90 degree angle to the stylus axis; with apical small projections. Promontory convex. - FEMALES: Measurements (n =12) DSW: 3.3-4.4; DSL: 4.0-5.4; CL: 1.3-1.6. FIVL: 4.9-5.6. ChL: 1.3-2.0 (Fig. 23D View Figure 23 ) DSS with constriction II well marked. Chelicerae similar to those of small males. Presence of a proapical spine on femur of pedipalpus. Femur and tibia IV unarmed. Tarsal segmentation: (n =12) 7, 10-11, 6-8, 7-8.


Similar to Ayacucho querococha sp. nov. because armed tibia IV (Fig. 7S, T View Figure 7 ). It differs from Ayacucho querococha sp. nov. by the set of the following characteristics: DSS with constriction II not-so-well marked (almost faint; Fig. 3C View Figure 3 ); a pair of spines on ocularium (Fig. 3C View Figure 3 ); coxa III with two apophyses (Fig. 3C View Figure 3 ; only one apophysis in Ayacucho querococha sp. nov.); femur IV without probasal row of tubercles; tibia IV without basal row of tubercles (Fig. 7S, T View Figure 7 ).


Roewer described Cargaruaya insignita in 1956 and Tapacochana insignita in 1957. Since they are both species in the genus Ayacucho , Tapacochana insignita Roewer, 1957 is a secondary homonym of Cargaruaya insignita Roewer, 1957 and must be replaced. Consequently, we create Ayacucho tapacocha nom. nov. as a replacement name for Roewer’s specific epithet.

Derivatio nominis.

The specific epithet, a noun in apposition, in reference to the type locality, Tapacocha, Ancash, Peru, as well as in relation to the name of the genus where the species was originally described by Roewer (1957).


(Fig. 28 View Figure 28 ) PERU. Ancash. Huascaran National Park; Tapacocha, Rio Fortaleza.

Material examined.

Type material: Holotype ♂, ' PERU, Ancash, Tapacocha | Rio Fortaleza , 3,200 m a.s.l., 04/III/1956, Weyrauch leg. (SMF RII 11644 View Materials /27) . Additional material: 2 ♂, 2 ♀ ' PERU, Ancash, Parque Nacional Huascarán, Chinancoche , 9°04′18″S 77°38′38″W | 15/V/2011, R. Pinto-da-Rocha & D. Silva leg. (MUSM) GoogleMaps ; 3 ♂, 3 ♀ ‘ditto’ (MUBI); 5 ♂, 5 ♀ ‘ditto’ (MZSP 36976). 2 ♂, 4 ♀ ' PERU, Ancash, Parque Nacional Huascarán, Laguna Paron , 8°59′43″S 77°40′41″W | 16/V/2011, R. Pinto-da-Rocha & D. Silva leg. (MUSM) GoogleMaps ; 2 ♂, 4 ♀ ‘ditto’ (MUBI); 4 ♂, 8 ♀ ‘ditto’ (MZSP 36977). 1 ♂, 2 ♀ ' PERU, Ancash, Parque Nacional Huascarán | Llaco , 9°28′11″S 77°27′50.8″W, 14/V/2011, R. Pinto-da-Rocha & D. Silva leg. (MUSM) GoogleMaps ; 2 ♂, 3 ♀ ‘ditto’ (MUBI); 3 ♂, 6 ♀ ‘ditto’ (MZSP 36978).














Ayacucho tapacocha

Benedetti, Alipio Rezende & Pinto-da-Rocha, Ricardo 2022

Tapacochana insignita

Benedetti & Pinto-da-Rocha 2022