Chelipoda modica Collin

Published, First, 2007, The Hemerodromiinae (Diptera: Empididae) of New Zealand II. Chelipoda Macquart, Zootaxa 1537 (1), pp. 1-88 : 12-13

publication ID

https://doi.org/ 10.11646/zootaxa.1537.1.1

persistent identifier

https://treatment.plazi.org/id/6A078784-BA6F-FFB1-AFFE-FAD41D99F3BA

treatment provided by

Felipe

scientific name

Chelipoda modica Collin
status

 

1. Chelipoda modica Collin View in CoL

[ Figs. 43–45 View FIGURES 40–48 , 74 View FIGURES 71–75 , 94 View FIGURES 91–103 , 133, 135 View FIGURES 131–139 , 143 View FIGURES 140–146 ]

Chelipoda modica Collin, 1928: 39–40 View in CoL .

Type material. Collin (1928) described Chelipoda modica from seven males and eight females from Ohakune, 12.xi.1922; iii., v–vii and ix–xi.1922 and 1923, TRH [ BMNH]. Ohakune is in: New Zealand, North Island, RI. A male in good condition was selected from Collin’s syntype series and is here designated lectotype with the label: Ohakune, 12.xi.1922, T. R. Harris [ BMNH]. The remaining 6♂, 8♀ labelled: Ohakune, iii., v–vii and ix–xi.1922 and 1923, T. R. Harris [ BMNH] from Collin’s syntype series are here designated paralectotypes.

Additional material: New Zealand, WO, TO, CO, HB, NC , KA, 29♂, 70♀ [ NMWC, NZAC and LUNZ] .

Description. Male: ( Fig. 143 View FIGURES 140–146 ), length 1.8–2.0mm (wet specimens).

Head: brown, thinly dusted grey, more strongly pale dusted on frons and face. Eyes virtually touching below antennae, widening below to mouth; ocl strong, black, diverging, slightly stronger than verticals which are arranged as an inner (vt1) and outer (vt2) pair; upo black, uniserial, diverging from ocular margin above becoming continuous with vt1; lpo pale, approximately biserial, very short; proboscis short, brownish; palpi short, pale with a single subapical seta. Antennae ( Fig. 94 View FIGURES 91–103 ) dark brown, basal segments slightly paler, sometimes yellowish; scape short and narrow bearing a distinct dorsal seta; pedicel wider, globular with apical circlet of small setae; postpedicel 1.5–2.0X as long as wide, about 0.4–0.5X length of arista.

Thorax: short; ground colour brown, slightly paler than head, sometimes appearing dark orange-brown in wet preserved specimens. brownish grey dusted dorsally with darker stripes which shift or disappear in different perspectives. Thoracic setae blackish brown; dc1 and dc4 absent, dc2 strong, dc3 slightly weaker and dc5 (prescutellar) weaker still; one pair of acr at front of mesonotum; sa and unp very strong, lnp much weaker; pprn and ph of similar length (about as strong as lnp); pa weak; one or two fine bristly hairs on pprnlb below pprn;

Legs: Yellow to brownish yellow, tarsi hardly darker, F 1 often contrastingly darker brown. C 1, only 0.8X length of thorax, armed sparsely with short pale bristles and hairs; C 2 and C 3 with short pale bristles anteroapically; F 1 about as long as C 1, strongly inflated above and below, about 2.8X as long as wide, widest 0.3 from base; femoral formula 5(4–6)/19(16–21)/11(9–13)/4(4–5), av and pv spines blackish, 1 or 2 basal bristles contiguous with line of av spines; posterior and dorsal faces of F 1 with fine hairs, a dorsal series of hairs being stronger and slightly outstanding. F 2 not especially slender, about 10X as long as wide; F 3 similar but slightly curved when viewed dorsally, a distinct dorsal ciliation of fine bristles longest in basal 0.6 where about as long as F 3 is deep. T 1 0.75X as long as F 1, only slightly curved, about 24 minute adpressed denticles ventrally behind which is a row of fine hairs and a few perpendicular cilia but otherwise short haired. T 2 and T 3 quite slender, short haired; at2 about 0.6–0.7X at3; pt1 = pt2 + pt3 + pt4.

Wing: ( Fig. 135 View FIGURES 131–139 ) extending considerably beyond tip of abdomen, veins brown, membrane faintly brown tinged; cell cup complete, rounded apically, vein A 1 strong throughout, continuing for a short distance beyond end of cell ( Fig. 133 View FIGURES 131–139 ); halteres white.

Abdomen: pale brown, hairs sparse and fine, rather stronger on posterior margin of tergite 5. Hypopygium ( Fig. 43, 44 View FIGURES 40–48 ) brown, rather strongly reflexed forwards; hypandrium and epandrium not fused; hypandrium large, rather keel-shaped, not divided posteriorly although less strongly sclerotized behind along median line. Epandrium divided, lobes narrowed apically and directed anteriorly; cerci small, fused with hypandrial lobes, bearing 3 short black spines on internal face and a ciliation of 4–5 short erect setae ( Fig. 45 View FIGURES 40–48 ). Lateral lobes on phallic process apically with a pair of small broadly triangular lateral processes. Postgonite (?) bearing 2 short blunt rod-like setae on inner face.

Description. Female: length 2.2–2.5mm (wet specimens).

Very similar to male, F 1 slightly stouter, only 2.6X as long as wide; femoral formula 5(4–6)/20(17–22)/ 12(10–13)/5(4–6). Sternite 8 with blunt point ( Fig. 74 View FIGURES 71–75 ); terminal papillae rather short with longish hairs.

Comments. A small, brownish species with C 1 pale, often yellowish and contrasting with the rest of the legs. Males are readily distinguishable from C. moderata by genitalic characters but determination of females can be problematical. Of a total of 114 specimens, most were captured in North Island forests in all months but July and August. The species is found in a variety of forest types from sea level to at least 1000 m. It can be found in small forest remnants surrounded by agricultural land as well as in large, intact forests. Only 4♂, 7♀ are known from the South Island, from two alpine sites in NC and FD. These are darker than North Island forms and perhaps reflect a tendency, widespread in New Zealand Chelipoda , for darker colouration in populations at higher altitudes. However, there appear to be small differences in the shape of the phallus and other genitalic characters but these were not considered to be of such significance as to warrant the erection of a separate species. There is some variation in the colour of the first antennal segment but there are 3♀ in NZAC from L. Ototoa [AK] leg. BAH, Poor Knights Is. [ND] leg RAC and Awanui, leg. MFT which have the segment clear yellow and probably represent an undescribed species allied with C. modica .

T

Tavera, Department of Geology and Geophysics

R

Departamento de Geologia, Universidad de Chile

NMWC

National Museum of Wales

NZAC

New Zealand Arthropod Collection

LUNZ

Lincoln University Entomology Research Museum

BAH

Biologische Anstalt Helgoland Marine Station

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Diptera

Family

Empididae

Genus

Chelipoda

Loc

Chelipoda modica Collin

Published, First 2007
2007
Loc

Chelipoda modica

Collin, J. E. 1928: 40
1928
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