Heraeus spinosus, Dellapé & Melo & Henry, 2016

Dellapé, Pablo M., Melo, María C. & Henry, Thomas J., 2016, A phylogenetic revision of the true bug genus Heraeus (Hemiptera: Rhyparochromidae: Myodochini), with the description of two new genera and 30 new species, Zoological Journal of the Linnean Society 177 (1), pp. 29-134 : 73-74

publication ID

https://doi.org/ 10.1111/zoj.12362

persistent identifier


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scientific name

Heraeus spinosus

sp. nov.


( FIGS 15H, 20M–P, 21B, 24)


Distiflagellomere with a wide sub-basal whitish band. Labium extending to mesocoxae. Profemur spotted, darker subapically, forming a band. Pygophore rounded, apex with a dorsally directed protuberance.

Description ( Fig. 15H)

Total length 6.27.

Head: Dark brown, shiny; with abundant short recumbent and long erect setae. Head length 1.34, width 0.96. Postocular length 0.48. Eyes not surpassing the dorsal margin of head in lateral view. Ocelli at level of an imaginary line passing the posterior border of eyes. Interocular width 0.48, interocellar width 0.24. Labium pale brown with long erect setae, almost extending to mesocoxae. Labial segment lengths: I 0.67, II 0.77, III 0.58, and IV 0.35. Antennae pale brown, except distal half of basiflagellomere and distiflagellomere darker, distiflagellomere with a wide sub-basal whitish band; all segments with abundant, short, semi-erect, and sparse erect setae. Antennal lengths: scape 0.55, pedicel 1.15, basiflagellomere 0.98, and distiflagellomere 1.13. Length of pale band on distiflagellomere 0.48.

Thorax: Anterior pronotal lobe dark brown, posteri- or lobe paler with irregular pale areas ( Fig. 15H), each humeral angle with a small whitish spot; collar delimited posteriorly by a punctate sulcus; anterior and posterior lobes with short recumbent and long erect setae, longest on anterior lobe. Pronotum pruinose, punctate, punctures larger on posterior lobe. Collar length 0.10, anterior lobe length 0.60, posterior lobe length 0.55; anterior lobe width 0.94, posterior lobe width 1.51. Pleurae brown, acetabular areas paler, punctate, with short recumbent setae. Evaporative area extended. Scutellum basally and a longitudinal median stripe dark brown, remainder paler brown, pruinose, punctate, with short recumbent and long erect setae, as on anterior pronotal lobe. Hemelytra brown, costal margin pale on proximal three-quarters, apical half of corium darker, with a subapical whitish spot and a small pale rounded spot internally ( Fig. 15H); setae short, semi-erect; membrane brown with a diffuse subapical pale spot, veins paler. Legs: Pale brown; profemur darker, mottled laterally and darker subapically forming a band, apex paler; coxae, protrochanter, a subapical band on meso- and metafemur, apex of tibiae, apex of tarsi, and pretarsi brown ( Fig. 21B); setae abundant and erect, longest on profemur. Profemur with spines restricted to apical two-thirds.

Abdomen: Brown, with abundant, short, recumbent setae. Male genitalia: Pygophore ( Fig. 20M, N) rounded, apex with a dorsally directed protuberance; slightly declivent posteriorly in lateral view. Anterior margin of dorsal aperture slightly rounded, inner projections quadrangular. Parameres: Figure 20 (O, P). Aedeagus: conjunctiva with spines laterally beneath the ejaculatory reservoir; vesica with a few spines laterally and two sclerotized lobes with a few spines distally; processus gonopori long and slender.


Brazil and Ecuador ( Fig. 24).


The specific epithet ‘ spinosus ’, Latin for spiny, refers to the spined aedeagus.

Type material

Holotype: ♂, BRAZIL: Rondonia, 62 km SW Ariquenes, near Fazenda Rancho Grande , 16/ 18-III-1996, BLT, U. Schmitz ( USNM) . Paratypes: 1♂, same locality as for holotype, 5–17-X- 1993, J.E. Eger, BLT ( USNM). BRAZIL : 1♂, Bahia: Encruzilhada, 960 m a.s.l., XI-1972, M. Alvarenga ( AMNH). ECUADOR: Napo : 1♀, vic. Puerto Misahuali, 1°2′4.2″S, 77°39′49.2″W, 1650–1900 ft, 6/ 19- IX-1998, mercury vapor and ultraviolet light, J.E. Eger ( USNM) GoogleMaps .


Smithsonian Institution, National Museum of Natural History


American Museum of Natural History













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