Clubiona filicata

Dankittipakul, Pakawin & Singtripop, Tippawan, 2008, Five new species of the spider genus Clubiona Latreille (Araneae: Clubionidae) from Thailand, Zootaxa 1747, pp. 34-60 : 37

Clubiona filicata O. P.- Cambridge, 1874

( Figs 5–7 , 30–33 )

Clubiona filicata O. P.- Cambridge, 1874: 413, fig. 35 (description of ɗ, Ψ); Gravely 1931: 261, fig. 16d; Tikader & Biswas 1981: 69, figs 120–121; Gong 1989: 109, figs 1–13; Majumder & Tikader 1991: 23, figs 30–35; Bis-

was & Raychaudhuri 1996: 199, figs 27–33; Song, Zhu & Chen 1999: 415, figs 245L–M, 248F–G. Clubiona swatowensis Strand, 1907: 562 (Description of Ψ); Strand 1909: 39, fig. 24.

Material examined. THAILAND: Chaiyaphum Province and District: 2ɗ, Thad Tone National Park, 15.– 20.xi.2006, leg. P. Dankittipakul ( MHNG, TNHM).

Diagnosis. Males of C. filicata can be easily recognized by the following features: an apical appendage of the male palp consists of a truncate embolus with a triangular retrolateral tubercle and a very long, filiform conductor that gives impression of the embolus ( Figs 6–7 , 30–32 ); a digitiform tegular apophysis ( Fig. 31 ) located anterio-prolaterally on the tegulum, situated close to the base of an apical appendage. Clubiona filicata closely resembles another species in the japonica -group, C. biembolata , in possessing a filiform conductor and enlarged posterior median eyes (see also Deeleman-Reinhold 2001: figs 67, 69). The latter species differs in the lack of a dorsal pattern on the opisthosoma (replaced by a lightly sclerotized dorsal scutum occupying ¾ of opisthosoma length) as well as the absence of a tegular apophysis on the male palp. The female can be recognized by its epigynal atrium situated anteriorly, by parallel insemination ducts running longitudinally to the posterior spermathecae and by the more or less triangular spermathecal heads.

Description. Male (Thad Tone National Park): Total length 4.2; prosoma 2.0 long, 1.4 wide; opisthosoma 2.2 long, 1.2 wide.

Coloration and pattern ( Fig. 5 ). Prosoma longly oval, narrowed in pars cephalica; in profile slightly higher behind longitudinal fovea; integument smooth, clothed with short fine hairs. Dorsal shield of prosoma pale yellow, slightly darker in front, without distinctive color pattern; fovea reddish. Chelicerae dark brown. Labium and endites yellowish brown. Sternum pale yellow. Legs pale yellow; anterior tibiae, metatarsi and tarsi distinctly darker. Opisthosoma elongate-oval; dorsum with a pattern consisting of dark purple blotches and stripes; venter pale. Spinnerets white.

Male palp ( Figs 6–7 , 30–33 ). Retrolateral tibial apophysis broad at base, gradually tapering towards its tip. Cymbium excavated dorso-apically, a profound depression covered with thick patch of short hairs. Bulb oval; sperm duct sinuate, forming a double loop. Tegular apophysis digitiform, situated proapically. Apical appendage consisting of truncate embolus and very long filiform conductor, forming a coiled structure, directed distad.

Natural history. The specimen examined was obtained from a Malaise trap set in a deciduous forest. The collecting locality, Thad Tone National Park, locates on Phu Lan Ka mountain range and consists of dipterocarp forest and lowland evergreen forest.

Distribution. South and Southeast Asia: India, Bangladesh, Pakistan, China and Thailand (new record, Fig. 69 ); probably common in tropical forests of Indochina.

Figures

FIGURE 69. Records of Clubiona species treated in the present study: * C. filicata; C. suthepica sp. n.; C. melanosticta; C. octoginta sp. n.; C. campylacantha sp. n.; Ο C. filifera sp. n.; C. abnormis sp. n.

FIGURES 5 – 10. Clubiona filicata O. P. - Cambridge, male from northeastern Thailand (5 – 7). 5 habitus, dorsal; 6 male palp, ventral; 7 ditto, retrolateral. Clubiona melanosticta Thorell, male from northern Thailand (8 – 10). 8 habitus, dorsal; 9 male palp, ventral; 10 ditto, retrolateral. Scale bars = 1.0 mm (5, 8); 0.1 mm (6 – 10).

FIGURES 30 – 33. Clubiona filicata O. P. - Cambridge, male from northeastern Thailand. 30 apical appendage of tegulum, ventral; 31 ditto, retrolateral; 32 ditto, prolateral; 33 retrolateral tibial apophysis.