Fannia terraba, Grisales & De Carvalho, 2019

Grisales, Diana & De Carvalho, Claudio J. B., 2019, Highland biodiversity of Fanniidae (Insecta, Diptera): fourteen new species from the Andes and Central America, Zootaxa 4551 (3), pp. 330-360 : 349-350

publication ID	https://doi.org/ 10.11646/zootaxa.4551.3.4
publication LSID	lsid:zoobank.org:pub:946C9A0F-D5C8-4EB6-8939-48BC7DE400A9
DOI	https://doi.org/10.5281/zenodo.5931488
persistent identifier	https://treatment.plazi.org/id/6909C857-F216-FF9B-5390-BF88FCFF3929
treatment provided by	Plazi
scientific name	Fannia terraba
status	sp. nov.

Treatment

Fannia terraba View in CoL sp. nov.

( Figs. 1L View FIGURE 1 , 4 View FIGURE 4 G–I, 5L, 7D, 8L)

Diagnosis. Fore tarsomeres 1–3 partially whitish–yellow, fifth fore tarsomere flattened, second fore tarsomere with setae forming a comb. Ventral katepisternal spine strong.

Description. Male holotype ( Fig. 1L View FIGURE 1 )

Head: eye setulose. Frontal vitta dark brown. 18 fr. 1 pair of oc, proclinate and developed, surrounded by long setulae. Fronto–orbital plate and parafacial with silver pollinosity, gena brown. Parafacial bare. Scape dark brown, 2 dorsal setae reaching pedicel. Pedicel black with golden yellow pollinosity, 1 long dorsal seta and surrounded by short setulae. Postpedicel dark brown, 2.5 times the length of pedicel. Arista dark brown, with short pubescence, rays shorter than width of base of arista. Palpus dark brown, claviform.

Thorax: dark brown; scutum without vittae. Acr 3:3, irregular rows. Dc 2:3. Pprn 3, developed, surrounded setulae. Pra 2, seta closest to suture longer. Proepimeron setulose. Katespisternum with long setulae and tuft of straight short setae ventrally.

Wing: brownish, with superior third, between C vein and superior half of R 2+3 vein darkened. Calypter whitish. Haltere brown.

Legs: coxae, femora and tibia brown; tarsi brown, except the 1–3 fore tarsomeres white yellowish, apex of the third tarsomere and pulvilli brownish. Fore femur on basal half with 1 row of av, apical half almost bare; 1 row of d; pd and p surface setulose, with thin and long hooked setae; 1 series of long pv. Fore tibia with 1 pre–apical d; 1 apical pv and 1 v. Fore second tarsomere on ventral surface with a tuft of short whitish setae ( Fig. 4G View FIGURE 4 ). Mid femur with 1 row of av, long on basal half and shorter on apical half; 1 row of ad on basal half; 1 row of unordered p, with 8 pre–apical differentiated setae; 1 row of pv with 2 basal longer setae; 1 row of short v, near pv row. Mid tibia strongly constricted on v surface on basal half and highly setulose on apical half ( Fig. 4H View FIGURE 4 ), 1 strong and long apical setae; 1 av apical; 2 ad on apical half; 1 sub–median and 1 pre–apical pd; 1 short pre–apical p; 1 pre–apical pv. Hind coxa with 1 seta on posterior margin and a tuft of long setae on anterior surface. Hind femur strongly curved and preapically enlarged, with long strong v and pv on the swelling ( Fig. 4I View FIGURE 4 ); 1 row of ad, 4 setae strongly differentiated on apical half; a long and strong, apical third bare; 2 av on the swelling, separated between them; tuft of long and strong v on the swelling, short setae on the median and basal third; 1 pre–apical d; pd surface with a short pre–apical seta; pv surface with a row of short setae; p surface bare on apical third. Hind tibia with 1 median and 1 pre–apical d; 1 sub–median ad; 2 sub–median and 1 apical av.

Abdomen: dark brown. Sternite 1 setulose. Sternite 5 ( Fig. 5L). Terminalia ( Fig. 7D View FIGURE 7 ): epandrium longer than wide, strong setae on basal half with two developed setae, two long and pointed apical internal projections; cercal plate much smaller than epandrium, with short and weak setae along the plate; surstylus fused with epandrium, curved, shaped as a comma in apical half, easily visible in frontal and lateral view, basal half with internal basal setae; bacilliform process absent.

Female: unknown.

Biology. Label information indicates that this species occurs in the highland forests of Costa Rica, 2100–3050 m. Nothing else is known about its habits or biology.

Comments. This species is similar to Fannia arcuata Chillcott, 1961 (pretiosa group) except it has dark legs. The new species is similar to Fannia gilvitarsis Chillcott, 1961 (pretiosa group) from which it differs in the paleyellow fore tarsus and apex of fifth tarsomere darkened.

Etymology. Name in apposition. The species epithet refers to the Terraba indigenous people, who once inhabited Puntarenas province in Costa Rica.

Type material. Holotype male ( INBio). COSTA RICA. Prov. Puntarenas, P.Int. / La Amistad, de Cerro Becón a Tres / Colinas, 2124m. 17 ABR 2002, M./ Alfaro , Libre, L _S_343750_565700/ #69143/ INBIO0003479937 INBIOCRI COSTA RICA . Paratypes. COSTA RICA. San José: Villa Mills, 3000m. Prov./ San Jose, Costa Rica / Curso Tachinidae y/ Syrphidae . 24 feb 1992 /L_S 390000, 495000/ INBIO CRI000 408439 (1 male, INBio); “ídem”/ INBIO CRI000 408387 (1 male, DZUP); “ídem”/ INBIO CRI000 408381 (1 male, INBio); “ídem”/ INBIO CRI000 408382 (1 male, INBio); Estac. Barva. Braulio/ Carrillo, N. P. 2500m / Heredia COSTA RICA / Feb 1990. G Rivera /233400–523200/ INBIO CRI000 202055 (1 male, INBio); “idem”/ Mar 1990. A Fernandez / INBIOCRI000 169464 (1 male, DZUP); Est. Cuerici, Sendero al Mirador, 4.6/km al E. de Villa Mills, San Jose / Costa Rica 2640–2700m. 20–22 ENE/1996. A Picado, de Luz /L_S 389700_499600 #6807/ INBIO CRI002 392887 (1 male, DZUP); Costa Rica, Prov. San Jose /Sendero a Cerro Cuerici, Limite del/ P.N. Chirripo. 3050m. 5 JUL 1996. A./ Picado. L _S_391300_500100 #7718/ INBIO CRI002 444089 (1 male, INBio)

The holotype is in good conditions.

Distribution. Costa Rica: Puntarenas province ( Fig. 8L View FIGURE 8 ).