Amatitlania kanna, Juan J. Schmitter-Soto, 2007

Juan J. Schmitter-Soto, 2007, A systematic revision of the genus Archocentrus (Perciformes: Cichlidae), with the description of two new genera and six new species., Zootaxa 1603, pp. 1-78 : 53-54

Amatitlania kanna , new species

Figures 17, 20

Archocentrus nigrofasciatus (part. et non Günther), Allgayer 1994: 12 (misidentification).

Cryptoheros cf. nigrofasciatus , Allgayer 2001: 18 (preliminary detection).

Holotype. FMNH 59243 , 83 mm SL (Fig. 20), E. H. Behre and J. Chambers, Feb. 2, 1923. San San swamp , Atlantic Panama.

Paratypes. BMNH 1925.3.6.119 (1), 1925.3.6.120-121 (2), 1925.3.6.122-123 (2), 1925.3.6.126 (1), FMNH 59240 (2), 59241 (1), 59242 (1), 116465 (1), MHNG 2646.78 (2), UMMZ 145716 (1).

Diagnosis. No unique autapomorphies, but differs from all other Amatitlania by secondary caudal pores absent (vs. present), peritoneum pigmented dorsolaterally (vs. uniformly), and quadrate bone wider than long (vs. not wider than long). Several features shared with Am. siquia (body relatively deeper, fewer circumpeduncular scales), some of them synapomorphies (Schmitter-Soto, in press): gill-rakers on first arch often bifid (vs. trapezoidal); caudal edge of urohyal sigmoid (vs. concave); anterodorsad spine on first dorsal pterygiophore present (vs. absent). However, it differs from its sister species as follows: dorsal-fin interradial scales distally in two rows (vs. one); posterior edge of mesethmoid rounded (vs. straight-irregular); gut simple (S- shaped, not folded anal and medial loops touching) (vs. shaped like an S folded ventrorostrally in adults, anal and medial loops not touching); and caudal blotch completely on fin (vs. partly on peduncle).

Description. D. XVII-XIX,8-10 (one specimen of 19 with 11 dorsal fin rays); A. IX-X,7-8. Gill rakers trapezoidal or bifid. Scales from lateral line to first dorsal-fin ray modally 2; circumpeduncular scales usually 16-17, modally 16; total vertebrae 27 (further meristic data appear in Table 3).

Largest specimen examined, 83 mm SL. The deepest-bodied Amatitlania , body depth 48-58% of SL, usually greater than 50% of SL (further morphometric data appear in Table 4). Head profile concave on orbits, convex on nape. Teeth conical, pointed, slightly retrorse. Upper symphysial teeth abruptly larger than adjacent teeth. Upper lip medially narrow; lower lip often with fleshy, hair-like papillae, not tapering nor squarish at corner, its lower angle acute.

Pectoral fins always reaching caudad beyond 2nd anal-fin spine pelvic fins always extending beyond 5th and often to 10th anal-fin spine. Filamentous rays of dorsal fin reaching to distal third of caudal fin. One or two pores continuing lateral line on caudal fin, no subsidiary scales. Dorsal-fin interradial scales arranged distally in one or two rows, up to 12 scales long; anal-fin scales usually in one row, up to 9 scales long.

Gut simple, usually slightly shorter than standard length. Peritoneum pigmented only dorsally, less so on upper sides. Genital papilla oval, but often distally expanded; uniformly pigmented on base.

Suborbital streak present; stripe from snout to eye usually well defined. Head bar interrupted in an area rostral to lateral line. Eyes blue with a golden rim. Bars on side of body sharp, medially and dorsally more intense; 4th bar I-shaped. No dots discernible on soft dorsal and anal fins. Breast olive. Axil of pectoral fin with same coloration as breast or dark, often with a dorsal spot; base of pectoral fin whitish. Caudal blotch on fin, across lateral line; often saddled.

Distribution. From rivers Cañaveral, Cricamola, Sixaola, and other localities in Atlantic Panama (Fig. 17).

Etymology. Greek καννα, meaning “a reed”, Río Cañaveral (= reedbed) being the first locality where the species was detected (see Remarks). A noun in apposition.

Remarks. The species was first recognized as distinct by Allgayer (2001: 18), who called his specimens from Rio Canaveral “ Cryptoheros cf. nigrofasciatus ”