Cybaeus coylei Bennett,

Bennett, Robb, Copley, Claudia & Copley, Darren, 2021, Cybaeus (Araneae: Cybaeidae): the aspenicolens species group of the Californian clade, Zootaxa 4926 (2), pp. 224-244: 233

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Cybaeus coylei Bennett

spec. nov.

Cybaeus coylei Bennett  spec. nov.

Figs 8–10View FIGURES 8–10, 13–15View FIGURES 11–13View FIGURES 14–19, 41View FIGURE 41

Type material. Holotype male. U.S.A.: California: Tuolumne County ¸ Pinecrest , 10.9.1959, W.J. Gertsch & V. D. Roth ( AMNH). 

Paratype. U.S.A.: California: Tuolumne. 1♀, Pinecrest , 10.9.1959, W.J. Gertsch & V  . D. Roth ( AMNH)  .

Etymology. The specific name is a patronym honouring arachnid ethologist and systematist Frederick A. Coyle, a valued friend and early mentor who guided Robb Bennett through his initial graduate work and with whom he experienced memorable road and field trips; name in genitive case.

Diagnosis. Cybaeus coylei  spec. nov. is grouped with C. aspenicolens  and C. blasbes  in the informal aspenicolens  subgroup. Distinguishing the aspenicolens  and fraxineus  subgroups is discussed in the aspenicolens  species group diagnosis. The male of C. coylei  spec. nov. is diagnosed by the relatively smoothly curved retrolateral margin of the massive, blunt patellar apophysis with about 2 dozen very small peg setae dorsally ( Figs 8View FIGURES 8–10, 13View FIGURES 11–13) as well as the strong dorsal deflection of the tip of the proximal arm of the tegular apophysis ( Fig. 10View FIGURES 8–10) such that much of the proximal arm is hidden in ventral view ( Fig. 9View FIGURES 8–10). The female of C. coylei  spec. nov. is diagnosed primarily by the form of the epigynum ( Fig. 14View FIGURES 14–19) which lacks any significant sclerotization and the relatively inconspicuous epigynal depression with a weakly defined anterior margin. Diagnostic differences distinguishing other species of the aspenicolens  subgroup from specimens of C. coylei  spec. nov. are discussed under C. aspenicolens  .

Description. Ventral tibia I macrosetae: 2–1p–2–1p– 1p.

Male: (n=1). About 40 or 50 stout peg setae retrolaterally on massive, blunt patellar apophysis; about two dozen very small peg setae dorsally ( Figs 8View FIGURES 8–10, 13View FIGURES 11–13). Proximal arm of tegular apophysis ( Figs 9–10View FIGURES 8–10) strongly deflected dorsally; large, broad, retrolaterally uncinate with tip weakly corkscrewed. Measurements. Holotype CL 3.3, CW 2.43, SL 1.55, SW 1.50.

Female: (n=1). Atrium ( Fig. 14View FIGURES 14–19) with paired unobscured openings; epigynal depression poorly developed, not strongly bordered anteriorly. Vulval ducts ( Fig. 15View FIGURES 14–19) contiguous from posterior end of copulatory ducts nearly to location of Bennett’s glands, apparently not joined anteriorly. Measurements. CL 3.4, CW 2.35, SL 1.55, SW 1.48.

Distribution. Known only from the type locality on the western flank of the southern Sierra Nevada in Tuolumne County ( Fig. 41View FIGURE 41).


Royal British Columbia Museum - Herbarium


American Museum of Natural History