Anabarhynchus iancommoni Ferguson

Ferguson, David J., Lambkin, Christine L. & Yeates, David K., 2014, Eight new species of Australian stiletto flies in the genus Anabarhynchus Macquart (Diptera: Therevidae) from South East Queensland, Zootaxa 3802 (4), pp. 553-582 : 560-564

Anabarhynchus iancommoni Ferguson sp. n.

( Figs 7 , 8 , 9 , 10 )

Type material. Holotype: Male. AUSTRALIA: Queensland: Yeppoon , I.F.B. Common, 18.xii.1964, (MEI_129408) ( ANIC _29:007865) ( ANIC). Condition: Micro-pinned dorsally into pith block.

Paratypes: 2 males. AUSTRALIA: Queensland: 2♂ same data as holotype; (MEI_129399) ( ANIC _29:007866); (MEI_129418) ( ANIC _29:007864) ( ANIC).

Diagnosis. Frons glossy black, scape length two-thirds length of the head; scutum with a pair of grey stripes separated by a blackish brown stripe; katepisternum with pile; post-spiracular pile absent; without hind coxal knobs, coxal macrosetae pale; all femora blackish brown; fore femur 1 pd, hind femur 1 av macroseta.

Description. Male. Body length: 7 mm. Wing length: 6 mm. ( Figs 7–10 ). Head. Lower frons raised, upper frons slightly concave, width 3.3 × ocellus width; ocellar tubercle flat; face and lower frons slightly protruding; parafacial viewed anteriorly bright grey, viewed either dorsally or ventrally appearing dark grey; lateral of antenna with irregular shaped blackish grey mark; lower frons predominantly with black pubescence, middle frons glossy black, upper frons matte black, with sparse grey pubescence towards eye margins; two rows of very short semierect black setae on upper half of frons; lower frons setae erect and restricted to area above antennal base, one fifth length of scape. Scape length two-thirds the length of the head measured along the lateral plane. Scape length 4.75 × pedicel length; scape and pedicel pale brown; f1 pale brown with numerous short dark setae on outer dorsolateral side and ventral basal quarter; f2 & f3 brown, covered with many small black setae, one-quarter the length of f1. Occiput with grey pubescence; several indistinct rows of black macrosetae, 25 setae on each side; postocciput to gena with grey pubescence, with long, pale, filiform pile; ventral corner of eye with distinctive swelling, numerous short black setae on ventral surface. Palp pale brown with pale hair-like setae; labellum pale brown; prementum with a few short black setae. Thorax. Integument brown-black; scutal chaetotaxy black (pairs): np, 3; sa, 2; pa, 1; dc, 5; sc, 2. Scutum with pair of grey vittae medially separated from each other by a blackish brown vittae of similar width. Pleura and coxae with thick pubescence, pleura when viewed dorsally are grey, ventrally black, posteriorly dark grey, anteriorly grey with a pair of shiny black bands on anterior surface of anepisternum/ katepisternum and anepimeron/meron. Katepisternum with pile dorsocentrally; prosternal furrow without pile; post-spiracular pile absent; coxae grey with long pile admixed with pale macrosetae; hind coxal knobs absent. Wing. Hyaline with pale brown infuscate, brown veined, stigma brown; costal setae beyond humeral cross-vein biserially arranged. Haltere. Pedicel basal half brown transitioning to pale yellowish apically, knob dorsally yellowish and ventrally dirty white. Legs. Fore femur 1 pd macroseta; hind femur 1 av macroseta; sparse, long, semi-erect pale pile on posterior surfaces of fore and middle femora, all femora blackish brown with short black setae; fore and hind tibia brown, middle femur yellow-brown; fore tarsi brown and middle and hind tarsi yellowbrown. Abdomen. Integument blackish brown; laterally compressed, tergites 2–7 with dorsal surfaces black, covered with short, appressed, dark setae; posterolaterally with bright grey pubescent marks, sparsely covered with erect pale pile laterally, tergites 2–3 when viewed anteriorly are bright grey while tergites 4–6 matte black, viewed posteriorly all tergites are matte black; tergite 2 apical band is bright white; sternites blackish brown, sternites 2–3 with grey pubescence sparsely covered with erect pale pile, sternite 4 with thin grey pubescent surface, with semierect dark pile, sternites 5–7 brown with semi-erect dark pile. Terminalia. (MEI_129399) (ANIC_29:007866): Epandrium ( Fig. 10 A): length 3/4 its width (measured along mid-line and between widest points when viewed dorsally); anterior margin broadly emarginate, greatly constricted towards the posterior, the apically margin deeply emarginate; sub-epandrial plate with numerous small black setae directed anteriorly. Gonocoxite ( Fig. 10 B): semispherical when viewed ventrally, black macrosetae on posterolateral margins; hypandrium translucent; outer gonocoxal process broadly triangular along the inner posteroventral edge and posteroventrally directed. Gonocoxal apodeme short. Inner gonocoxal process short, reduced in size in comparison to the gonostylus, ventrally directed, with weak apical setae. Gonostylus posteriorly directed, ventral edge ridged with long dark setae along inner edge, lateral surface broad, dorsal edge with laterally directed lobe medially, apex truncated and curved dorsally. Ventral lobe rounded and indistinct. Aedeagus ( Figs 10 C, D): distiphallus curved ventrally; parameral sheath pale and broadly triangular; ventral apodeme broad with a flattened bilobed apex; lateral ejaculatory apodeme narrow, bandlike; ejaculatory apodeme laterally compressed medially, with dorsal ridge.

Variation. Male. Frons width 3.3–3.4 × ocellus; scape length: 4.2–4.5 × width; occipital setae 22–28 each side.

Female. Unknown.

Etymology. The specific epithet is derived to honor the late Dr. Ian. F.B. Common, entomologist and collector of the type specimens.

Comments. Known from three males collected in late December at Yeppoon , Queensland.

Anabarhynchus iancommoni sp. n. keys to A. argenteus Lyneborg in couplet 43 in Lyneborg (2001), with both species possessing pile on the katepisternum. Anabarhynchus iancommoni sp. n. can be readily separated from A. argenteus because A. iancommoni sp. n. has antennal length longer than the head when measured along the lateral plane, post-spiracular pile absent, and with pale coxal macrosetae. Anabarhynchus argenteus has antenna shorter than the head, post-spiracular pile present, and black coxal macrosetae.

Anabarhynchus iancommoni sp. n. is very similar to A. oblongicornus Winterton with both having many small black setae on the dorsolateral surfaces of f1, f2 & f3 ( Fig. 9 ); post-spiracular pile absent and lacking hind coxal knobs, which are secondary reduced in A. kampmeierae Irwin & Lyneborg. Anabarhynchus iancommoni sp. n. can be separated from A. oblongicornus , with A. iancommoni sp. n. having blackish brown femora, scape length twothirds the head length measured along the lateral plane, and with some hair-like pile on the dorsocentral area of the katepisternum. Anabarhynchus oblongicornus has yellow-brown femora, a scape length equal to head length, and is without pile on the katepisternum. Anabarhynchus iancommoni sp. n. appears to be most closely aligned to species in Lyneborg’s maritimus species-group.

Figures

FIGURE 7. Anabarhynchus iancommoni sp. n., holotype male (MEI _ 129408) (ANIC _ 29: 007865): lateral view. Scale line = 2 mm.

FIGURE 8. Anabarhynchus iancommoni sp. n., holotype male, (MEI _ 129408) (ANIC _ 29: 007865): oblique view. Scale line = 2 mm.

FIGURE 9. Anabarhynchus iancommoni sp. n., close-up of antenna, holotype male, (MEI _ 129408) (ANIC _ 29: 007865): Scale line = 1 mm.

FIGURE 10. Anabarhynchus iancommoni sp. n., paratype male terminalia (MEI _ 129399) (ANIC _ 29: 007866): A, epandrium, left dorsal view, right ventral view; B, gonocoxite left dorsal view, right ventral view; C, aedeagus, dorsal view; D, aedeagus, lateral view; Scale line = 0.5 mm.